The Ecozones ofthe World

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Jürgen Schultz The Ecozones ofthe World The Ecological Divisions ofthe Geosphere Translated into English by Bridget Ahnert Second Edition with 144 Figures, 22 Tables, and 5 Boxes 4ü Springer

Contents General: the treatment ofthe ecozones and global overviews of selected characteristics 1 Distribution 7 2 Climate 11 2.1 Solar radiation 11 2.2 The growing season and conditions for plant growth 14 3 Relief and drainage 17 3.1 Geomorphological processes 17 3.2 Drainage and water balance 17 4 Soils 21 4.1 Soil fertility 21 4.2 Soil water budget 23 4.3 Soilunits and soil zones 26 5 Vegetation and animals 35 5.1 Structural characteristics of the Vegetation 35 5.2 Ecosystem model of ecozone 38 5.3 Available supply of organic matter in the ecosystem 40 5.4 Primary production 40 5.4.1 Photosynthesis and respiration 40 5.4.2 Primary production from plant Stands 42 5.4.3 Production capacity of the world's plant cover 43 5.5 Consumption by animals and secondary production 48 5.6 Waste and decomposition 49 5.7 Turnover ofminerals 51 6 Land use 55

VIII Contents Regional section: The individual ecozones 7 Polar subpolar zone 63 7.1 Distribution 63 7.2 Climate 64 7.2.1 Temperature, length of day, precipitation 64 7.2.2 Annual temperature changes in the soil and air layer next to the soil 65 7.2.3 Summer solar radiation and heat budget 67 7.3 Relief and drainage in periglacial areas 68 7.4 Soils 72 7.5 Vegetation and fauna in the tundra and polar deserts 73 7.5.1 Distribution of Vegetation 74 7.5.2 Biomass and primary production 75 7.5.3 Animals and animal feed 76 7.5.4 Decomposition and turnover of minerals 77 7.5.5 Model of a tundra ecosystem 78 7.6 Landuse 80 Synoptic diagram for the tundra 81 8 Boreal zone 83 8.1 Distribution 83 8.2 Climate 84 8.3 Relief and drainage 87 8.4 Soils 89 8.5 Vegetation and animals 91 8.5.1 Boreal coniferous forest 91 8.5.2 Peatbogs 92 8.5.3 Forest tundra, polar and forest tree lines 93 8.5.4 Biomass and primary production 94 8.5.5 Decomposition, organic soil matter and mineral reserves 94 8.5.6 Boreal coniferous forest ecosystems 98 8.6 Landuse 98 Synoptic diagram for the Boreal zone 100 9 Temperate midlatitudes 103 9.1 Distribution 103 9.2 Climate 103 9.3 Relief and drainage 105 9.4 Soils 107 9.5 Vegetation and animals 108 9.5.1 Seasonality in deciduous forests 109 9.5.2 Water budget in forests 111 9.5.3 Biomass and primary production, growth and litter production 112

Contents IX 9.5.4 Mineral budget in midlatitude deciduous broadleaf and boreal coniferous forests 113 9.5.5 Model of an ecosystem of a deciduous forest 118 9.6 Landuse 118 Synoptic diagram for the Temperate midlatitudes 121 10 Dry midlatitudes 123 10.1 Distribution 123 10.2 Climate 124 10.3 Relief and drainage 125 10.4 Soils in the steppes 125 10.4.1 Types of soil in the zone 125 10.4.2 Halomorphic soils 128 10.5 Vegetation and animals in the steppes 128 10.5.1 Types of steppes 129 10.5.2 Life-form and adaptation to winter cold and summer drought 130 10.5.3 Animals 130 10.5.4 Biomass, primary production and decomposition 131 10.5.5 Availablesupplyandturnoverofrninerals 134 10.6 Landuse 134 10.6.1 Large scale grain cultivation 134 10.6.2 Extensive pasture economy 136 Synoptic diagram for steppes 138 11 Subtropics with winter rain 141 11.1 Distribution 141 11.2 Climate 142 11.3 Relief and drainage 143 11.4 Soils 143 11.5 Vegetation and animals 144 11.5.1 Sclerophyllous Vegetation 144 11.5.2 Adaptation to summer drought 146 11.5.3 Animals 148 11.5.4 Fire 148 11.5.5 Biomass and primary production 150 11.6 LandUse 152 Synoptic diagram for the Subtropics with winter rain 154 12 Subtropics with year-round rain 157 12.1 Distribution 157 12.2 Climate 158 12.3 Relief and drainage 159 12.4 Soils 160 12.5 Vegetation 161 12.5.1 Structural characteristics 161

X Contents 12.5.2 Available supply and turnover in a semi-evergreen oak forest in the southeastern United States 162 12.6 Landuse 165 Synoptic diagram for the Subtropics with year-round rain 165 13 Dry tropics and subtropics 169 13.1 Distribution 169 13.2 Climate 170 13.3 Relief and drainage 172 13.3.1 Weathering processes andcrusts 172 13.3.2 Aeolian processes 172 13.3.3 Stream erosion andwash denudation 173 13.4 Soils 174 13.5 Vegetation and animals 175 13.5.1 Vegetation and soil water budget 176 13.5.2 Adaptation to drought and stress 180 13.5.3 Animals in the desert 183 13.5.4 Biomass and primary production 184 13.6 Landuse 185 13.6.1 Pastoral nomadism 185 13.6.2 Oasis agriculture 187 Synoptic diagram for tropical thorn savannas and subtropical thorn... 189 steppes in the semi-arid Dry tropics and subtropics 189 14 Tropics with summer rain 193 14.1 Distribution 193 14.2 Climate 194 14.3 Relief and drainage 195 14.4 Soils 198 14.4.1 General characteristics of soils in the subtropics and tropics with summer and year-round rain 198 14.4.2 Soil types 200 14.5 Vegetation and animals 202 14.5.1 Structural characteristics of savanna Vegetation 202 14.5.2 Animals 203 14.5.3 Savanna fires 204 14.5.4 Biomass and primary production 204 14.5.5 Zoomass and animal feed 206 14.5.6 Decomposition of litter 208 14.5.7 Mineral supplies and turnovers 209 14.6 Landuse 210 Synoptic diagram for the Tropics with summer rain 213 15 Tropics with year-round rain 215 15.1 Distribution 215 15.2 Climate 216

Contents XI 15.3 Relief and drainage 217 15.4 Soils 218 15.5 Vegetation and animals 221 15.5.1 Structure ofthe tropical rainforest 221 15.5.2 Dynamics ofthe Vegetation 224 15.5.3 Animals 224 15.5.4 Biomass and primary production 225 15.5.5 Animal feed 226 15.5.6 Litter, the litter layer, decomposition and humus 226 15.5.7 Mineral supplies and turnovers 227 15.5.8 Rainforest ecosystem 229 15.6 Landuse 230 Synoptic diagram for the Tropics with year-round rain 235 References 237