Giorgio MANZI Paleoanthropologist



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Giorgio MANZI Paleoanthropologist # Professional address : Sapienza Università di Roma, Dipartimento di Biologia Ambientale, Piazzale Aldo Moro 5, 00185 Roma (Italy) # Telephone : (39) 06 4991 2271 (office) / 2690 (lab) / 2495 (fax) # Email : giorgio.manzi@uniroma1.it # Web : https://sites.google.com/a/uniroma1.it/giorgio-manzi # Birth place/date : Rome, February 9th 1958 # Nationality/status : Italian, married # University degree: Doctor in Biological Sciences # Present position : Associate Professor, Sapienza Università di Roma # Present appointments : Director of the Museum of Anthropology «Giuseppe Sergi», Roma Director of the Polo museale Sapienza for the years 2013-2015 and 2016-2018 # Past eminent appointments : Vice-secretary of the Istituto Italiano di Antropologia (IsItA) and Assistant editor of the Rivista di Antropologia (Journal of Anthropological Sciences) for the period 1991-2003 General Secretary of the Istituto Italiano di Paleontologia Umana (IsIPU) for the years 1999-2006 Member of the Permanent Council of the International Association for the Study of Human Paleontology 1

# Grants & prices : Research grants credited by the Italian Ministry of Instruction (MIUR/urst), Ministry of Culture (MIBAC), Research National Council (CNR) and by the Sapienza Università di Roma P.I. of the Awards attributed by the Sapienza University of Rome for the year 2013 - prot. C26H13A45J Award Fabio Frassetto for Physical Anthropology in 2006, credited by the Accademia Nazionale dei Lincei # Research activity : Author or co-author of papers in scientific journals, which include: Nature Communications, Proceedings of the National Academy of Sciences USA, The Anatomical Record, Evolutionary Anthropology, Journal of Human Evolution, Quaternary Science Reviews, American Journal of Physical Anthropology, PLoS One, American Journal of Human Biology, Current Anthropology, Journal of Anthropological Sciences etc. Editor or contributor of monographic volumes. Main research interests are in human paleontology and skeletal biology, including in particular: i) studies on fossil specimens from the Early, Middle and Late Pleistocene, as well as the Holocene, from Italy, Spain, Libya and Ethiopia; ii) special focus on topics such as the earliest human presence in Europe, origin and fate of the Neanderthals, evo-devo interpretation of the fossil record with special reference to the origin of Homo sapiens; iii) researches concerning human populations of central Sahara during the Holocene; iv) projects involving the study of human skeletal samples from Italy dated from the Neolithic to the Early Middle Ages. Methodological approaches include morphology with either traditional or multivariate approaches, geometric morphometrics and treatment of CT data; phenetic evaluation of numerical data: interpretation of the epigenetic discrete traits of the human cranium, with inferences about ontogeny in extinct hominines. Other interests are in dental anthropology, paleodemography, paleopathology, forensic anthropology, and history of science. # Field experience : Field activities in various prehistoric sites in Italy and abroad, including Grotta Breuil (Monte Circeo, Italy; Late Pleistocene), various sites in the Tadrart Acacus massif and surroundings (Fezzan, Libya), Sima the los Huesos cave site (Sierra de Atapuerca, Spain; Middle Pleistocene), Lower Paleolithic localities in central Italy (co- Director of the excavation campaigns 2000-2006 at Ceprano-Campogrande and 2007-2010 at Colle della Pece), Melka Kunture localities (Ethiopia), Laetoli and Olduvai (Tanzania). # Editorial work & conferences : Member of scientific or organizing committees of congresses, symposia, and colloquia; among the most recent: Humans in the focus of our evolution (16-19 March 2005, Roma-Pofi-Cerpano, Italy); Nostro fratello Neanderthal: when we were not alone (Circeo National Park, Sabaudia, Italy). Invited speaker in a number of international congresses (Johannesburg, Gibraltar, Zagreb, Tempe ). Member of the Comité de Lecture of the «Bulletin and Mémoires de la Société d'anthropologie de Paris»; since 2001. Associate Editor of the «American Journal of Physical Anthropology»; since 2012. Member of the Editorial Board of the «Rendiconti Lincei - Scienze Fisiche e Naturali», Accademia Naz. Lincei; since 2014. Reviewer for various scientific journals, including «American Journal of Human Biology», «American Journal of Physical Anthropology», «Bulletin and Mémoires de la Société d'anthropologie de Paris», «Comptes-rendus de 2

l'académie des Sciences de Paris», «Comptes Rendus Palevol», «Current Anthropology», «Journal of Anthropological Sciences» (già «Rivista di Antropologia»), «Journal of Archaeological Sciences», «Journal of Human Evolution», «Nature», «Proceedings of the National Academy of Sciences USA». # Teaching : Teaching activity at the Sapienza Università di Roma. Present courses for the Faculty of Natural Sciences: Paleoanthropology, Human Ecology and Natural History of the Primates, Museology; for the School of Specialization in Archaeology: human paleontology Member of the Faculty of PhD programs in Biologia Animale (1999-2010) and in Biologia Ambientale and Evoluzionistica (since 2011) of the Sapienza Università di Roma Tutorial assistance in MA, MSc, PhD theses. Other current teaching activities: member of the scientific committee and lecturer for the Master-school of Paleoanthropology held in Perugia, Italy; years 2011-2016. # Museology & dissemination : Active since 1983 with different roles in the Museum of Anthropology «Giuseppe Sergi» of the Sapienza Università di Roma; since 2004 he is the Director of the same museum; since 2012 he is also the Director of the Polo museale Sapienza (center of coordination of the 20 museums of the Sapienza Università di Roma). Contributed on different levels to various exhibitions. Among the most recent are worth of mention: the collaboration to the concept, planning and realisation of "La scimmia nuda" at the Museo Tridentino di Scienze Naturali (April 2007 - January 2008) with following transfers; the role of consultant for Paleoanthropology at the exhibition "Homo Sapiens": la grande storia della diversità umana" at the Palazzo delle Esposizioni in Rome (November 2011 - April 2012) with following transfer. Author of a number of dissemination papers (in newspapers, magazines, websites etc.); participant/curator in/of TV as well as radio programs. Guest-editor of special issues of newspapers and magazines. Several conferences and seminars in Italy (on average 15/year in the last years); also in Spain, France, and Switzerland. Author of popular books, published by Il Mulino (Bologna, Italy): Homo sapiens (2006), L evoluzione umana (2007), Uomini e ambienti (2009, co-authored with A. Vienna), Scimmie (2011, co-authored with J. Rizzo), Il grande racconto dell evoluzione umana (2013); published by Laterza (Roma-Bari, Italy): La scienza delle nostre origini (2013, coauthored with C. Tuniz and D. Caramelli), translated in the USA as The Science of Human Origins (Left Coast Press, 2014). # Bibliometric data : Scopus, 11/01/2016 h-index 19 documents 42 citations 927 (by 621 documents) 3

# Selected references 200-2014 : 1. Ascenzi A., Mallegni F., Manzi G., Segre A.G. & Segre Naldini E. 2000. A re-appraisal of Ceprano calvaria affinities with Homo erectus, after the new reconstruction. Journal of Human Evolution 39: 443-450. 2. Bastir M., Rosas A., Gunz Ph., Peña-Melián A., Manzi G., Harvati K., Kruszynski R., Stringer C. & Hublin J.J. 2011. Brain base evolution in highly encephalised human species. Nature Communications (DOI: 10.1038/ncomms1593). 3. Bruner E. & Manzi G. 2001. Allometric analysis of the skull in Pan and Gorilla by geometric morphometrics. Rivista di Antropologia 79: 45-52. 4. Bruner E. & Manzi G. 2002. The virtual replica of Nazlet Khater, Egypt. Cranium and mandible: first results. In P.M. Vermeersch (ed.), Paleolithic Quarrying Sites in Upper and Middle Egypt; pp. 337-345. Egyptian Prehistory Monographs - Leuven University Press, Leuven. 5. Bruner E. & Manzi G. 2003. Towards a re-appraisal of the Early Neolithic skeleton from Lama dei Peligni (Abruzzo, Italy). Computed tomography and 3D reconstruction of the cranium. Rivista di Antropologia 81: 69-78. 6. Bruner E. & Manzi G. 2004. Variability in facial size and shape among North and East African human populations. Italian Journal of Zoology 71: 51-56. 7. Bruner E. & Manzi G. 2005. CT-based description and phyletic evaluation of the archaic human calvarium from Ceprano, Italy. The Anatomical Record (part A), 285A: 643-658. 8. Bruner E. & Manzi G. 2006. Saccopastore 1: the earliest Neanderthal? A new look at an old cranium. In K. Harvati & T. Harrison (eds), Neanderthals Revisited: New Approaches and Perspectives; pp. 23-36. Springer, Dordrecht. 9. Bruner E. & Manzi G. 2007. Landmark-based shape analysis of the archaic Homo calvarium from Ceprano (Italy). American Journal of Physical Anthropology, 132: 355-366. 10. Bruner E. & Manzi G. 2008. Paleoneurology of an early Neanderthal: endocranial size, shape, and features of Saccopastore 1. Journal of Human Evolution, 54: 729-742. 11. Bruner E., Ricci F. & Manzi G. 2002. Faces from the ancient Fezzan: a geometric morphometric approach. In S. Di Lernia & G. Manzi (eds), Sand, Stones, and Bones. The archaeology of death in the Wadi Tanezzuft Valley (5000-2000 bp); pp. 251-260. The Archaeology of Libyan Sahara, vol. I (AZA Monographs 3) - All'Insegna del Giglio, Firenze. 12. Bruner E., Averini M. & Manzi G. 2003. Endocranial traits. Prevalence and distribution in a recent human population. European Journal of Anatomy 7: 23-33. 13. Bruner E., Manzi G. & Arsuaga J.L. 2003. Encephalisation and allometric trajectories in the genus Homo. Evidence from the Neandertal and modern lineages. Proceedings of the National Academy of Sciences USA 100: 15335-15340. 14. Bruner E., Saracino B., Ricci F., Tafuri M., Passarello P. & Manzi G. 2004. Midsagittal cranial shape variation in the genus Homo by geometric morphometrics. Collegium Antropologicum 28: 99-112. 15. Bruner E., Mantini S., Perna A., Maffei C. & Manzi G. 2005. Fractal dimension of the middle meningeal vessels: variation and evolution in Homo erectus, Neanderthals, and modern humans. European Journal of Morphology, 42: 217-224. 16. Bruner E., Manzi G. & Holloway R. 2006. Krapina and Saccopastore: endocranial morphology in the pre- Würmian Europeans. Periodicum Biologorum, 108: 433-441. 4

17. Dal Poz M., Ricci F., Reale B., Malvone M., Salvadei L. & Manzi G. 2001. Paleobiologia della popolazione altomedioevale di San Lorenzo di Quingentole, Mantova. Cranio e scheletro postcraniale. In A. Manicardi (a cura di), San Lorenzo di Quingentole. Archeologia, Storia ed Antropologia; pp. 151-198. Società Archeologica Padana (S.A.P. srl), Mantova. 18. Di Lernia S. & Manzi G. (eds) 2002. Sand, Stones, and Bones. The archaeology of death in the Wadi Tanezzuft Valley (5000-2000 bp). All'Insegna del Giglio (AZA Monographs 3), Firenze. 19. Di Lernia S., Bertolani G.B., Merighi F., Ricci F.R., Manzi G. & Cremaschi M. 2001. Megalithic architecture and funerary practices in the late prehistory of Wadi Tanezzuft (Libyan Sahara). Libyan Studies 32: 29-48. 20. Di Vincenzo, F. & Manzi G. 2013. Social learning and origin of the language faculty by means of natural selection. Journal of Anthropological Sciences, 91: 23-30. 21. Di Vincenzo F., Churchill S.E. & Manzi G. 2012. The Vindija Neanderthal scapular glenoid fossa: a comparative shape analysis. Journal of Human Evolution, 62: 274-285. 22. Manzi G. 2001. The earliest diffusion of the genus Homo toward Asia and Europe: A brief overview. In P.V. Tobias, M.A. Raath, J. Moggi-Cecchi, G.A. Doyle (eds), Humanity from African Naissance to Coming Millenia; pp. 117-124. Firenze University Press - Witwatersrand University Press, Firenze - Johannesburg. 23. Manzi G. 2003. Epigenetic cranial traits, Neandertals and the origin of Homo sapiens. Rivista di Antropologia 81: 57-67. 24. Manzi G. 2004.Human evolution at the Matuyama-Brunhes boundary.evolutionary Anthropology13:11 24. 25. Manzi G. 2004. Italian prehistoric promenades: the human fossil sample. From the earliest European to the latest Neadertals. In Baquedano E. & Rubio Jara S. (eds), Miscelánea en Homenaje a Emiliano Aguirre, vol. III; pp. 220-230. Museo Archeológico Regional, Alcalá de Henares. 26. Manzi G. (ed.) 2005. The Ceprano hominid: collection of reprints in occasion of the international conference "Humans in the Focus of our Evolution". Università 'La Sapienza', Roma. 27. Manzi G. 2011. Before the emergence of Homo sapiens: Overview on the Early-to-Middle Pleistocene fossil record (with a proposal about Homo heidelbergensis at the subspecific level). International Journal of Evolutionary Biology, vol. 2011, ID 582678: 1-11 (doi: 10.4061/2011/582678). 28. Manzi G. 2012. On the trail of the genus Homo between archaic and derived morphologies. Journal of Anthropological Sciences, 90: 99-116. 29. Manzi G. & Di Vincenzo F. 2012. Le dernier ancêtre de l homme moderne. Pour La Science 411: 20-27. 30. Manzi G. & Di Vincenzo F. 2013. Light has been thrown (on HumanOrigins). A Brief History of Paleoanthropology, with'notes on the Punctuated Origin of Homo sapiens. Aisthesis, VI (2): 31-48. 31. Manzi G. & Ricci F. 2003. Populations of the Roman Era in the central Sahara: Skeletal samples from the Fezzan (south-western Libya) in a diachronic perspective. In M. Liverani (ed.), Arid Lands in Roman Times, AZA Monographs 4; pp. 15-22. All'Insegna del Giglio, Firenze. 32. Manzi G., Gracia A. & Arsuaga J-L. 2000. Cranial discrete traits in the Middle Pleistocene humans from Sima de los Huesos (Sierra de Atapuerca, Spain). Does hypostosis represent any increase in "ontogenetic stress" along the Neandertal lineage? Journal of Human Evolution 38: 425-446. 33. Manzi G., Bruner E., Caprasecca S., Gualdi G. & Passarello P. 2001. CT-scanning and virtual reproduction of the Saccopastore Neanderthal crania. Rivista di Antropologia 79: 61-72. 34. Manzi G., Mallegni F. & Ascenzi A. 2001. A cranium for the earliest Europeans: Phylogenetic position of the hominid from Ceprano, Italy. Proceedings of the National Academy of Sciences USA 98: 10011-10016. 5

35. Manzi G., Ricci F. & Salvadei L. 2002. Porotic hyperostosis as marker of health and nutritional conditions: Response to B. Rothschild. American Journal of Human Biology 14: 417-420. 36. Manzi G., Bruner E. & Passarello P. 2003. The one-million-year-old Homo cranium from Bouri (Ethiopia): a reconsideration of its H. erectus affinities. Journal of Human Evolution 44: 731-736. 37. Manzi G., Magri D., Milli S., Palombo M.R., Margari V., Celiberti V., Barbieri M., Barbieri Mz., Melis R.T., Rubini M., Ruffo M., Saracino B., Tzedakis P.C., Zarattini A. & Biddittu I. 2010. The new chronology of the Ceprano calvarium (Italy). Journal of Human Evolution, 59: 580-585. 38. Manzi G., Micheli M., Di Vincenzo F., Caramelli D. & Piperno M. 2010. La porzione di scapola dell Uomo di Altamura: estrazione, documentazione e analisi morfologica. DIRE in Puglia (M.B.A.C., Direzione Regionale per i Beni Culturali e Paesaggistici della Puglia), 2: 41-46. 39. Manzi G., Magri D. & Palombo M.R. 2011. Early Middle Pleistocene environmental changes and human evolution in the Italian peninsula. Quaternary Science Reviews, 30: 1420-1438. 40. Marangoni A., Belli M.L., Caramelli D., Moggi-Cecchi J., Zavattaro M. & Manzi G. 2011., Tierra del Fuego, its ancient inhabitants, and the collections of skeletal remains in the Museums of Anthropology of Florence and Rome. Museologia Scientifica n.s., 5 (1-2): 88-96. 41. Marangoni A., Caramelli D. & Manzi G. 2014. Homo sapiens in the Americas. Overview of the earliest human expansion in the New World. Journal of Anthropological Sciences, 92: 79-97. 42. Martinón-Torres M., Bermúdez de Castro J.M., Gómez-Robles A., Arsuaga J.L., Carbonell E., Lordkipanidze D., Manzi G. & Margvelashvili A. 2007. Dental evidence on the hominin dispersals during the Pleistocene. Proceedings of the National Academy of Sciences USA, 104: 13279-13282. 43. Minozzi S., Manzi G., Ricci F., Di Lernia S., & Borgognini Tarli S.M. 2003. Nonalimentary tooth use in prehistory: An example from Early Holocene in Central Sahara (Uan Muhuggiag, Tadrart Acacus, Libya). American Journal Physical Anthropology 120: 225-232. 44. Mounier A, Condemi S, Manzi G. 2011. The stem species of our species: a place for the archaic human cranium from Ceprano, Italy. PLoS one, 6(4), e18821: 1-11 (doi: 10.1371/journal.pone.0018821). 45. Muttoni M., Scardia G., Kent D.V., Swisher C.C., Manzi G. 2009. Pleistocene magnetochronology of early hominin sites at Ceprano and Fontana Ranuccio, Italy. Earth and Planetary Science Letters, 286: 255 268. 46. Ricci F., Manzi G., Fornai C., Vecchi F. & Passarello P. 2002. The human skeletal remains: inventory and inferences. In S. Di Lernia & G. Manzi (eds), Sand, Stones, and Bones. The archaeology of death in the Wadi Tanezzuft Valley (5000-2000 bp); pp. 217-250. The Archaeology of Libyan Sahara, vol. I (AZA Monographs 3) - All'Insegna del Giglio, Firenze. 47. Ricci F., Fornai C., Tiesler Blos V., Rickards O., di Lernia S. & Manzi G. 2008. Evidence of artificial cranial deformation from the Late Prehistory of South-Western Libya. International Journal of Osteoarchaeology, 18: 372-391. 48. Salvadei L., Ricci F. & Manzi G. 2001. Porotic hyperostosis as marker of health and nutritional conditions during childhood. Studies at the transition between Imperial Rome and the Early Middle Ages. American Journal of Human Biology 13: 709-717. 49. Tafuri M., Mastroroberto M. & Manzi G. 2003. Human skeletal remains from the Middle Bronze Age cemetery of Sant'Abbondio (Pompeii, Italy). Rivista di Antropologia 81: 79-107. 50. Tafuri M.A., Bentley R.A., Manzi G., Di Lernia S. 2006. Mobility and kinship in the prehistoric Sahara: strontium isotope analysis of Holocene human skeletons from the Acacus Mts. (southwestern Libya). Journal of Anthropological Archaeology, 25: 390-402. 6