Curriculum Vitae for Ronni N Glud

Curriculum Vitae for Ronni N Glud 1 Born: Nationality: Family: Current work address: 10. Nov.1963 Danish Married, 3 children Scottish Association of Marine Science Dunstaffnage Marine Laboratory Oban, Argyll PA37 1QA Phone: +44 (0) 1631 559427 E-mail: ronnie.glud@sams.ac.uk Scientific education & positions: 1990; Master in Microbial Ecology, Aarhus University, DK 1993; Ph.D. in Microbial Ecology, Aarhus University, DK 1993-1996; Assistant Professor, Max Planck Institute-Bremen, D 1997; Assistant Professor Aarhus University, DK 1997-2007; Associated Professor, University of Copenhagen, DK 2007; Dr. Scient at Faculty of Science, University of Copenhagen, DK 2007-present; Professor in Marine Biogeochemistry at SAMS, UK (2010; Professor in Marine Biogeochemistry at Southern Danish University) Research Interest: Development and application of in situ lander technology for studying microbial ecology and biogeochemistry. Benthic biogeochemistry and solute exchange, carbon mineralization, photosynthesis, bioturbation, bioirrigtaion. Development and application of microsensors and imaging techniques in microbial ecology and biogeochemistry. Polar biology, climate changes, local regional and global element cycling, sea-ice microbiology. Grants that have been rewarded as PI (or Co-PI) - last 5 years: Glud (2005) National Science Foundation-Denmark Carbon, nitrogen and microbial dynamics in permeable sands - Total amount 143.000 Dkr Glud and Eyre (2005) Fellowship from Southern Cross University Nitrogen cycling in permeable sand Total amount 80.000 Glud and Middelboe (2006-2009) Natural Science Foundation Denmark, Benthic microbial processes on a microscale: Importance of viruses II. Total amount 600.000 Dkr Gribsholt and Glud, (2006-2009) Natural Science Foundation Denmark Patterns in fauna behavior and activity. Total amount 2.300.000 Dkr Thamdrup et al. (2006-2009) Natural Science Foundation Denmark, Galathea 3 oxygen minimum zones. Partner amount 300.000 Dkr

Glud (2007-2008) Natural Science Foundation Denmark, An ultra-deep diving lander for hadal studies, Total amount 650.000 Dkr Gissel-Nielsen, Rysgaard et al (2007-2010) Royal Scientific Investigations Greenland Ecosystem West Greenland, Partner amount 50.000 Dkr Sejr et al (2007-2010) Royal Scientific Investigations Greenland Ecosystem metabolism and CO2 exchange in Arctic fjord-systems, Partner amount, 200.000 Dkr Eyre & Glud (2007-2012) Australian Research Council Unravelling the pathways of nitrogen cycling in permeable sandy sediments in tropical ecosystems, Partner amount, 200.000 Dkr Glud, Stahl, Solan (2008-2011) NERC Carbon mineralization of shelf and coastal sediments: A holistic approach using state of the art lander technology and eddy correlation technique Total amount 3.200.000 Dkr Davison & Glud (2009-2011) NERC Microheterogeneity of carbon mineralization and metal mobilization in marine sediments Partner amount 2.800.000 Dkr Partner in EU project HYPROX (2009-2011) Partner amount 1.900.000 Dkr Turnewitsch et al (2009-2012) NERC Impact of kilometer-scale seafloor topography on deep sea biogeochemistry Partner amount 1.400.000 Dkr Glud (2010-2011) FNU The Arctic marine carbon cycle: Consequences of changes in climate and sea-ice cover Partner amount 2.800.000 Dkr International Scientific Activities - Last 5 years: Cruises & expeditions 2005- Expedition to NE Greenland 7-month sabbatical at Southern Cross University (Australia) 2006 2 x Cruises in Sagami Bay, Japan Research visit to West Greenland (Natur instituttet - Nuuk) 2007 Galathea cruise off Chile-Peru Research visit to West Greenland (Natur instituttet - Nuuk) 2008 Cruise in Sagami Bay, Japan 3 x Research visit to West Greenland (Natur instituttet - Nuuk) Cruise off South west Greenland 2009 Sea-ice camp in the central Arctic Ocean Research visit to West Greenland (Natur instituttet - Nuuk) Research visit to Singapore (Danish Hydraulic Institute) Expedition to Heron Island, Australia 2

Active participation/first author contributions at larger International Scientific meetings last 5 years: 3 2005 Summer meeting of America Society of Limnology and Oceanography, Santiago de Compostella, Spain Invited speaker Gordon Research Conference on Chemical Ocenogr, Tilton, NH, USA invited discussion leader 2006 National Sea-Research Conference, Denmark, Odense, Denmark Invited keynote speaker 2008 Nereis-Park Bioturbation meeting, Renesse, The Netherlands, - Invited keynote speaker ISSM, International symposium for subsurface microbiology Shizuoka, Japan,- Invited speaker (tutorial) 2009 ASLO, Nice, France Invited speaker (tutorial) AGU-cryosphere meeting, Nuuk, Greenland Invited speaker (3 invited seminars GEOMAR, D; University of Virginia, US Stony Brook University, US)

Esteem: Associate editor for Limnology and Oceanography Chair of Science committee for Scottish Marine Institute Board member of Greenland Ecosystem Monitoring Program (GEM) 4 Student - supervision: Finalized Master-projects N. Munksby (University of Copenhagen) - Benthic fauna and mineralization K. Hancke (University of Copenhagen)- Benthic diatoms and Carbon cycling G. a Nordi (University of Copenhagen) - Microniches in marine sediments M. Hansen (University of Copenhagen) - The effect on Thyrasia on C and S cycling M. Sturlurson (University of Copenhagen) - Microbial carbon degradation in the pelagic L Carlsson (University of Copenhagen) - Benthic viruses M Jørgensen (University of Copenhagen) - Benthic viruses DM Mikkelsen (University of Copenhagen) - Sea-ice microalgae succession in West Greenland M Kristensen (University of Copenhagen) Gas dynamics in Sea-ice A Soegaard (University of Copenhagen) - ph and salinity effects on sea-ice microalgae M Larsen (University of Copenhagen) - N and O 2 dynamics in Amphiura filiformis communities Ph.D. Projects: O. Greef (MPI Bremen, Germany) Title: Entwicklung eines benthischen Landers zur Bestimmung von in situ Sulfatreduktionsraten. W. Rieß (MPI -Bremen, Germany) Title: Benthic exchange processes of nutrients, O2, CO2: Laboratory and in situ studies. F. Wenzhöfer (MPI- Bremen, Germany) Title: Biogeochemische Prozesse beim Abbau Organischer Substanz in Marinen Sedimenten, CO2 Freisetzung. Ongoing Ph.D. Projects: G a Nordii (Fiskeri Hoejskolen, Thorshavn) Effect of aqua-culture on benthic processes M Blicher (University of Copenhagen) Benthic infauna and their importance for the function of marine sediments in Arctic M Carlsson (Southern Danish University) Aquaculture, carbon and nitrogen cycling

M Larsen (SAMS) Microscale dynamics of marine sediments 5 G Turner (SAMS) Benthic O2 turn-over studied by eddy correlation Catja Carreira (University of Copenhagen) Virus microscale dynamics D Soegaard (GN Nuuk) Sea-ice microbiology and associated gas exchange

6 Publication list Published International peer-reviewed Journals and books only: (Citations >2800; recent years >350 citations yr -1, H-factor 32) 1. RN Glud, NB Ramsing, NP Revsbech (1992) Photosynthesis and photosynthesiscoupled respiration in natural bio-films quantified with microsensors. J Phycol 28: 51-60 2. RN Glud, JK Gundersen, NP Revsbech, BB Jørgensen (1993) In Situ measurements of total and diffusive oxygen uptake of sediments pp 427-430. In: Trends in Microbial Ecology (ed. R. Guerrero and C. Pedros-Alio) Proc. 6th In. Symp. Microbial Ecol. 3. DE Canfield, BB Jørgensen, H Fossing, RN Glud, JK Gundersen, NB Ramsing, B Thamdrup, JW Hansen, LP Nielsen, POJ Hall (1993) Pathways of organic carbon oxidation in three continental margin sediments. Mar Geol 113: 27-40 4. RN Glud, JK Gundersen, NP Revsbech, BB Jørgensen (1994) Effects on the benthic diffusive boundary layer imposed by microelectrodes. Limnol Oceanogr 39: 462-467 5. B Thamdrup, RN Glud, JW Hansen (1994) Manganese oxidation and in situ manganese fluxes from a coastal sediment. Geochim Cosmochim Acta 58: 2563-2570 6. RN Glud, JK Gundersen, BB Jørgensen, NP Revsbech, HD Schulz (1994) Diffusive and total oxygen uptake of deep-sea sediments in the eastern South Atlantic Ocean: In Situ and laboratory measurements. Deep Sea Res 41: 1767-1788 7. RN Glud, K Jensen, NP Revsbech (1995) Diffusivity in surficial sediments and benthic mats determined by use of a combined N2O-O2 microsensor. Geochim Cosmochim 59: 231-237 8. RN Glud, JK Gundersen, NP Revsbech, BB Jørgensen, M Hüttel (1995). Calibration and performance of the stirred flux chamber from the benthic lander Elinor. Deep Sea Res 42: 1029-1042 9. J Lorenzen, RN Glud, NP Revsbech (1995). Impact of microsensor caused changes in diffusive boundary layer thickness on O2 profiles and photosynthetic rates in benthic communities of microorganisms. Mar Ecol Prog Ser 119: 237-241 10. A Tengberg, De Bovee F, Hall P, Berelson W, Cicceri G, Crassous P, Devol A, Emerson S, Glud RN,Graziottin F, Gundersen J K, Hammond D, Helder W, Holby O, Jahnke R, Khripounoff A, Nuppenau V, Pfannkuche O, Reimers C, Rowe G, Sahami A, Sayles F, Schuster M, Wehrli B, and De Wilde P (1995) Benthic chamber and profile landers in oceanography - A review of design, technical solutions and functioning. Prog in Oceanogr 35: 253-294 11. H Fossing, Gallardo VA, Jørgensen BB, Hüttel M, Nielsen LP Schulz H, Canfield DE, Forster S, Glud RN, Gundersen JK, Küver J, Ramsing NB, Teske A, Thamdrup B,

Ulloa O (1995) Concentration and transport of nitrate by the mat-forming sulphur bacterium Thioploca. Nature 374: 713-715 7 12. LP Nielsen and RN Glud (1996) Denitrification in a coastal sediment measured in situ by nitrogen isotope pairing technique applied to a benthic flux chamber. Mar Ecol Prog Ser 137: 181-186 13. M Kühl, RN Glud, H. Ploug, N.B. Ramsing (1996) Microenvironmental control of photosynthesis and photosynthesis-coupled respiration in an epilithic cyanobacterial biofilm. J Phycol 32: 799-812 14. B. Thamdrup, D.E. Canfield, T.G. Ferdelman, RN Glud, J.K. Gundersen (1996) A biogeochemical survey of the anoxic basin Golfo Dulce, Costa Rica. Revista de Biologia Tropical 44: 19-33 15. RN Glud, NB Ramsing, JK Gundersen, I Klimant (1996) Planar optrodes, a new tool for fine scale measurements of two dimensional O2 distribution in benthic communities. Mar Ecol Prog Ser 140: 217-226 16. RN Glud, S Forster, M Hüttel (1996) Influence of radial pressure gradients on solute exchange in stirred benthic chambers. Mar Ecol Prog Ser 141: 303-311 17. G Holst, RN Glud, M Kühl, I Klimant (1997) A microoptrode array for fine scale measurements of oxygen distribution. Sensors and Actuators B 39: 122-129 18. I Klimant, M Kuhl, RN Glud, G Holst (1997) Optical measurements of oxygen and other environmental parameters in microscale: Strategies and biological applications. Sensors and Actuators B 38-39: 29-37 19. C Hensen, H Landenberger, M Zabel, JK Gundersen, RN Glud, HD Shulz (1997) Simulation of early diagenetic processes in continental slope sediments off Southwest Africa: The computer model CoTam tested. Mar Geol 144: 191-210 20. JK Gundersen, NB Ramsing, RN Glud (1998) Predicting the signal of oxygen microsensors from physical dimensions, temperature, salinity, and oxygen concentrations. Limnol Oceanogr 43: 1932-1937 21. C Lassen, RN Glud, NB Ramsing, NP Revsbech (1998) A method to improve the spatial resolution of photosynthetic rates obtained by oxygen microsensors. J phycology 34: 89-93 22. RN Glud, CM Santegoeds, D de Beer, O Kohls, NB Ramsing (1998) Oxygen dynamics at the base of a biofilm studied with planar optrodes. Aqua Microb Ecol 14: 223-233 23. O Greeff, RN Glud, O Holby, JK Gundersen, BB Jørgensen (1998) A benthic lander for tracer studies in the sea bed: In situ measurements of sulfate reduction. Continent Shelf Res 18: 1581-1594 24. RN Glud, O Holby, F Hofmann, DE Canfield (1998) Benthic mineralisation in Arctic sediments (Svalbard). Mar Ecol Prog Ser 173: 237-251

8 25. T Fenchel, RN Glud (1998) Veil architecture in a sulphide-oxidizing bacterium enhances countercurrent flux. Nature 394: 367-369 26. RN Glud, I Klimant, G Holst, O Kohls, V Meyer, M Kühl, JK Gundersen (1999) Adaptation, test and in situ measurements with O2 microoptodes on benthic landers. Deep Sea Res 26: 171-183 27. S Forster, RN Glud, JK Gundersen, M Huettel (1999) In situ study of bromide tracer and oxygen flux in coastal sediments. Estu, Coast and shelf Science 49:813-827 28. JE Kostka, B Thamdrup, RN Glud, DE Canfield (1999) Rates and parthways of carbon oxidation in permanently cold Arctic sediments. Mar Ecol Prog Ser 180: 7-21 29. RN Glud, M Kühl, O Kohls, NB Ramsing (1999) Heterogenity of oxygen production and consumption in a photosynthetic microbial mat as studied by planar optodes. J Phycol 35:270-279 30. RN Glud, JK Gundersen, O Holby (1999) Benthic In situ respiration in the upwelling area off central Chile. Mar Ecol Prog Ser 186:9-18 31. RN Glud, T Fenchel (1999) The importance of ciliates for interstitial solute transport in benthic communities. Mar Ecol Prog Ser 186:87-93 32. C Reimers, RN Glud (2000) In situ chemical sensor measurements at the sedimentwater interface. In Chemical sensors in Oceanography Gordon and Breach Science Publishers (ed. M Varney) Chapter 12: 249-282 33. PJ Hansen, A Skovgaard, RN Glud DK Stoecker (2000) Physiology of the mixotrophic dinoflagellate Fragilidium subglobosum. II Effects of time scale and prey concentration on photosynthetic performance. Mar Ecol Prog Ser 201:137-146 34. F Wenzhöfer, O Holby, RN Glud, HK Nielsen, JK Gundersen (2000) In situ microsensor studies of a hydrothermal vent at Milos (Greece). Mar Chem 69:43-54 35. RN Glud, JK Gundersen, NB Ramsing (2000) Electrochemical and optical oxygen microsensors for in situ measurements. In in situ monitoring of aquatic systems: Chemical analysis and speciation. John Wiley & Sons Ltd (eds J Buffle & G Horvai). Chapter 2:19-73 36. RN Glud, Risgaard-Petersen N, Thamdrup B, Fossing H, Rysgaard S (2000) Benthic carbon mineralization in a high-arctic sound. Mar Ecol Prog Ser 206:59-71 37. T Fenchel, RN Glud (2000) Benthic Primary production and O 2 -CO 2 dynamics in a shallow water sediment: Spatial and temporal heterogeneity. Ophelia 53:159-171 38. B König, G Holst, O Kohls, T Richter, RN Glud, M Kühl (2001) Imaging of oxygen distribution at benthic interfaces: a brief review. In: Organism-Sediment Interactions,. Eds.: J.Y.Aller, S.A.Woodin & R.C.Aller. Univ of South Carolina Press, pp 63-73.

9 39. S Rysgaard, M Kühl, RN Glud, JW Hansen (2001) Biomass, production and horizontal patchiness of sea ice algae in a high-arctic fjord (Young Sound, NE Greenland). Mar Ecol Prog Ser 223:15-26 40. M Kühl, RN Glud, J Borum, R Roberts, S Rysgaard (2001) Photosynthetic performance of surface associated algae below sea ice as measured with a pulse amplitude modulated (PAM) fluorometer and O 2 microsensors. Mar Ecol Prog Ser 223:1-14 41. RN Glud, A Tengberg, M Kühl, POJ Hall, I Klimant, G Holst (2001) An in situ instrument for planar O2 optode measurements at benthic interfaces. Limnol and Oceanogr 46:2073-2080 42. RN Glud, JK Gundersen (2002) Exchange and microdistribution of solutes at the benthic interface: an in situ study in Aarhus Bight, Denmark. In: Environmental Electrochemistry: Analyses of Trace Element Biogeochemistry. American Chemical Society. (M. Taillefert Ed). Chapter 8: 144-161 43. R Roberts, M Kühl, RN Glud, S Rysgaard (2002) Primary production of coralline red algae in a high Arctic Fjord. J Phycol 38:273-283 44. RN Glud, S Rysgaard, M Kühl (2002) A laboratory study on O 2 dynamics and photosynthesis in sea ice algal communities: Quantification by microsensors, O 2 exchange rates, 14 C-incubations and PAM fluorometry. Aqua Microb Ecol 27:301-311 45. F Wenzhöfer, RN Glud (2002) Benthic carbon mineralisation in the Atlantic: A synthesis based on in situ data from the last decade. Deep-Sea Res 49:1255-1279 46. RN Glud, M Kühl, F Wenzhöfer, S Rysgaard (2002) Benthic diatoms of a high arctic fjord (Young Sound NE Greenland): Importance for ecosystem primary production. Mar Ecol Prog Ser 238:15-29 47. RN Glud, N Blackburn (2002) The effect of chamber size on benthic oxygen uptake measurements: A simulation study. Ophelia 56:23-31 48. N Munksby, M Benthien, RN Glud (2002) Flow induced flushing of relict burrow structures in central Skagerrak (Norway). Marine Biology 141:939-945 49. PB Christensen, RN Glud, T Dalsgaard, P Gillespie (2003) Implications of long line mussel farming on benthos, oxygen and nitrogen dynamics in coastal sediments. Aquaculture: 218:567-588 50. RN Glud, JK Gundersen, H Røy, BB Jørgensen (2003) Seasonal dynamics of benthic O 2 uptake in a semi enclosed bay: Importance of diffusion and fauna activity. Limnol Oceanogr 48: 1265-1276 51. M Middelboe, RN Glud, K Finster (2003) Distribution of viruses and bacteria in relation to diagenetic activity in an estuarine sediment. Limnol Oceanogr 48: 1447-1456 52. Riisgard HU, O Kinne, T Fenchel, E Fee, R Hesslein, J Elser M Scranton, J Cole, N Hairston, E Canuel, RN Glud, C Nielsen, P Beninger, R Waagbo, GI Hemre, G Hulata, T

10 Hoisaeter, D McLusky, R Kneib, S Shumway, R Warwick, L Hagerman, PS Larsen, J Ott, PC Dworschak, F Boero, JM Gili K Philippart, M Seaman (2003) Misuse of the peer-review system: Time for countermeasures Mar Ecol Prog Ser 258: 297-309 53. S Rysgaard, RN Glud, (2004) Anaerobic N 2 production in Arctic sea-ice. Limnol Oceanogr 49:86-94 54. RN Glud, S Rysgaard, T Fenchel, PH Nielsen (2004) A conspicuous H 2 S oxidizing microbial mat from a high-latitude Arctic fjord (Young Sound, NE Greenland). Mar Biol 145: 51-60 55. S Rysggard, RN Glud, N Risgaard-Petersen, T Dalsgaard (2004) Denitrification and anammox activity in Arctic sediments. Limnol Oceanogr 49: 1493-1502 56. F Wenzhöfer, RN Glud (2004) Small-scale spatial and temporal variability in benthic O 2 dynamics of coastal sediments: Impact of fauna activity. Limnol Oceanogr 49: 1471-1481 57. RN Glud, M Middelboe (2004) Virus and bacteria dynamics of a coastal sediment: implication for benthic carbon cycling. Limnol Oceanogr 49:2073-2081 58. K Hancke, RN Glud (2004) temperature effects on respiration and photosynthesis in three diatom dominated benthic communities. Aqua Microb Ecol 37:265-281 59. A Tengberg, P Hall, U Andersson, B Lidén, O Styrenius, G Boland, F de Bovee, B Carlsson, S ceradini, A Devol, G Duineveld, JU Friemann, RN Glud, A Khripounoff, J Leather, P Linke, L Lund-Hansen, G Rowe, P santschi, P de Wilde, U Witte (2005) Intercalibration of benthic flux chambers II. Hydrodynamic characterization and flux comparison of 14 different designs. Mar Chem 94:147-173 60. BB Jørgensen, RN Glud, O Holby (2005) Oxygen distribution and bioirrigation in Arctic fjord sediments. Mar Ecol Prog Ser 292:85-95 61. SI Jensen, M Kühl, RN Glud, LB Jørgensen, A Priemé (2005) Oxygen microzones and radial oxygen loss from roots of Zostera marina. Mar Ecol Prog Ser 293:49-58 62. RN Glud, F Wenzhöfer, A Tengberg, M Middelboe, K Oguri, H Kitazato (2005) Distribution of oxygen in surface sediments from central Sagami Bay, Japan: In situ measurements by microelectrodes and planar optodes. Deep Sea Res I 52:1974-1987 63. B Koenig, O Kohls, G Holst, RN Glud, M Kühl (2005) Fabrication and test of sol-gel based planar optodes for use in aquatic sediments. Mar Chem 97:262-276 64. MS Frederiksen, RN Glud (2006) Oxygen dynamics in the rhizosphere of Zostera marina: A two-dimensional planar optode study. Limnol Oceanogr 51:1072-1083 65. Oguri K, H Kitazato, RN Glud (2006) Platinum octaetylporphyrin based planar optodes combined with UV-LED excitation light source: an ideal tool for high resolution O 2 imaging in O 2 depleted environments. Mar Chem 100:95-107

11 66. M Middelboe, RN Glud, F Wenzhöfer, K Oguri, H Kitazato (2006) Spatial distribution and activity of viruses in the deep-sea sediments of Sagami Bay, Japan. Deep Sea Res I 53:1-13 67. M Middelboe, RN Glud (2006) Viral activity along a trophic gradient in continental margin sediments off central Chile. Mar Biol Res 2:41-51 68. RN Glud (2006) Microscale techniques to measure photosynthesis: A mini-review (p31-41). In Functioning of microphytobenthos (eds J Kromkamp, FC Brouwer, GF Blanchard, RM Forster, V Creach). Proceedings from Amsterdam-colloquium 21-23 August 2003, pp 31-42. 69. PLM Cook, F Wenzhöfer, S Rysgaard, OS Galaktinov, FJR Meysman, BD Eyre, J Cornwell, M Huettel & RN Glud (2006) Quantification of denitrification in permeable sediments: Insight from 2 dimensional simulation analysis and experimental data. Limnol Oceanogr Method 4:294-307 70. TG Ferdelman, B Thamdrup, DE Canfield, RN Glud, J Kuever, R Lillebæk, NB Ramsing, C Wawer (2006) Biogeochemical controls on the oxygen, nitrogen and sulfur distributions in the water column of Golfo Dulche, Costa Rica: an anoxic basin revisited. Revista de Biologia Tropical 54:171-191 71. H Stahl, A Glud, CR Schröder, I Klimant, A Tengberg, RN Glud (2006) Time-resolved ph imaging in marine sediments with a luminescent planar optode. Limnol Oceanogr Method 4:336-345 72. K Timmermann, GT Banta, RN Glud (2006) Linking Arenicola marina irrigation behaviour to oxygen transport and dynamics in sandy sediments. J Mar Res 64:915-938 73. G Roeselers, TB Norris, RW Castenholz, S Rysgaard, RN Glud, M Kühl, G Muyzer (2007) Diversity of phototrophic bacteria in microbial mats from Arctic hot springs (Greenland). Environ Microbiol 9:26-38 74. RN Glud, P Berg, H Fossing, BB Jørgensen (2007) Effects of the diffusive boundary Layer (DBL) on benthic mineralization and O 2 distribution: A theoretical model analysis. Limnol Oceanogr 52:547-557 75. RN Glud, S Rysgaard, M Kühl, JW Hansen (2007) The sea-ice in Young Sound: Implications for C Cycling In: Carbon cycling in Arctic marine Ecosystems: Case study Young Sound. Meddr Greenland, Bioscience Special issue. (S. Rysgaard & RN Glud Eds). Chapter 4, 58:62-85 76. B Thamdrup, RN Glud, JW Hansen (2007) Benthic carbon cycling in Young Sound, NE Greenland. In: Carbon cycling in Arctic marine Ecosystems: Case study Young Sound. Meddr Greenland, Bioscience Special issue. (S. Rysgaard & RN Glud Eds). Chapter 8, 58:138-157 77. RN Glud, S Rysgaard (2007). The annual organic carbon budget of Young Sound, NE Greenland. In: Carbon cycling in Arctic marine Ecosystems: Case study Young Sound.

Meddr Greenland, Bioscience Special issue. (S. Rysgaard & RN Glud Eds). Chapter 11, 58:194-203 12 78. S Rysgaard, RN Glud (2007). Carbon cycling and climate change: Predictions for a high-arctic marine ecosystem (Young Sound, NE Greenland). In: Carbon cycling in Arctic marine Ecosystems: Case study Young Sound. Meddr Greenland, Bioscience Special issue. (S. Rysgaard & RN Glud Eds). Chapter 12, 58:206-213 79. JW Behrens, HJ Stahl, JF Steffensen, RN Glud (2007) Oxygen dynamics around buried lesser sandeel, Ammodytes tobiannus (Linnaeus, 1785); mode of ventilation and metabolic requirements. Journal of Experimental Biology 210:1006-1014 80. S Rysgaard, RN Glud, MK Sejr, J Bendtsen, PB Christensen (2007) Inorganic carbon transport during sea-ice growth and decay: A carbon pump in the polar seas. J Geophys Res Ocean. 112: CO3016 81. PLM Cook, F Wenzhöfer, F Janssen, M Hüttel, RN Glud, (2007) Benthic solute exhange and carbon mineralization in two shallow subtidal sandy sediments: Impact of advective porewater exchange. Limnol Oceanogr 52:1943-1963 82. S. Tankere-Müller, H Zang, W Davidson, N Finke, O Larsen, H Stahl, RN Glud (2007) Fine scale remobilisation of Fe, Mn, Co, Ni, Cu and Cd in contaminated marine sediment. Mar Chem 106:192-207 83. BD Eyre, RN Glud, N Patten (2008) Mass coral spawning a natural large scale nutrient addition experiment. Limnol Oceanogr 53: 997-1013 84. RN Glud, BD Eyre, N Patten (2008) Biogeochemical responses to coral mass spawning at the Great barrier Reef: Effects on respiration and primary production. Limnol Oceanogr 53: 1014-1024 85. NL Patten, Mitchell JG, Middelboe M, Eyre B, Seuront L, Harrison PL, Glud RN (2008) Bacterial and viral dynamics during a mass spawning period on the Great Barrier Reef. Aqua Microb Ecol 50:209-220 86. S Rysgaard, RN Glud, MK Sejr, ME Blicher, HJ Stahl (2008) Denitrification activity and oxygen dynamics in Arctic sea-ice. Polar Biol 31:527-537 87. K Hancke, TB Hancke, LM Olsen, G Johnsen, RN Glud (2008) Temperature effects on microalgae photosynthesis-light responses measured by O 2 -production, pulse amplitude modulation (PAM) fluorescence and 14C-assimilation. J Phycol 44:501-514 88. RN Glud (2008) Oxygen dynamics of marine sediments. Mar Biol Res 4:243-289 89. MS Jørgensen, RN Glud, M Middelboe (2008) Virus dynamics in a coastal sediment: Seasonal pattern, controlling factors and relations to the pelagic-benthic coupling. Mar Biol Res 3:165-179

90. DM Mikkelsen, S Rysgaard, RN Glud (2008) Microalgal composition and primary production in Arctic sea ice a seasonal study from Kobberfjord/ Kangerluarsunnguaq, West Greenland. Mar Ecol Prog Ser 368:65-74 13 91. RN Glud, H Stahl, P Berg, F Wenzhofer, K Oguri, H Kitazato (2009) In situ microscale variation in distribution and consumption of O 2 : A case study from a deep ocean margin sediment (Sagami Bay, Japan). Limnol Oceanogr 54:1-12 92. RN Glud, B Thamdrup, H Stahl, F Wenzhoefer, A Glud, H Nomaki, K Oguri, NP Revsbech, H Kitazato (2009) Nitrogen cycling in a deep ocean margin sediment (Sagami bay, Japan). Limnol Oceanogr 54:723-734 93. S Rysgaard, J Bendtsen, LT Pedersen, H Ramløv, RN Glud (2009) Increased CO 2 uptake due to sea-ice growth and decay in the Nordic Seas. J Geophys Res 114: C09011 94. P Berg, RN Glud, A Hume, H Stahl, K Oguri, V Meyer, H Kitazato (2009) Eddy correlation measurements of oxygen uptake in deep ocean sediments. Limnol Oceanogr Meth 7:576-584 95. NP Revsbech, RN Glud (2009) Biosensor for laboratory and lander-based analysis of benthic nitrate plus nitrite distribution in marine environments. Limnol. Oceanogr Meth 7:761-770 96. RN Glud, J Woelfel, U Karsten, M Kuhl, S Rysgaard (2009) Benthic microalgal production in the Arctic: Status of the current database. Bot Mar 52:559-571 97. MS Carlsson, RN Glud, JK Petersen (submitted) Degradation of mussel (Mytilus edulis) fecal pellets released from hanging long-lines upon sinking and after settling at the sediment. Can J Fish Aqua Science 98. OS Galaktinonov, FJR Meysman, RN Glud, JJ Middelburg (submitted) Oxygen penetration around burrows and roots in aquatic sediments. J Mar Res 99. DH Søgaard, M Kristensen, S Rysgaard, RN Glud, PJ Hansen, KM Hillingsøe (submitted) Autotrophic and heterotrophic activity in Arctic first-year sea-ice: Seasonal study from Marlene Bight, SW Greenland. Mar Ecol Prog Ser

Selected popular presentation (News papers, TV radio etc): My research has on a number of occasions been described in public media and magazines. The article list below only include presentations were I have directly involved as co-author and not contributions where I only have contributed with an interview or press release. News papers, magazines, popular books PB Christensen, S Rysgaard, RN Glud (2000) Ishavet et varmt emne. Artikel i Dagbladet information PB Christensen, S Rysgaard, RN Glud (2000) Iltfrit havis fjerner kvælstof. Artikel i Dagbladet information PB Christensen, S Rysgaard, RN Glud (2000) Alger en overset faktor. Artikel i Dagbladet information T Fenchel, RN Glud (2002) Iltsvind og debatforurening. JyllandsPosten - kronik RN Glud (2005 ) Marine eutrophication and benthic metabolism chapter 12 in an internet textbook (Eds P Wassmann & K Olli) M Middelboe, RN Glud (2006). Virus- havbundens mindste beboere. Naturens Verden 89:32-40. RN Glud, M Kühl (2006) Havbundens stofomsætning I: Havet (Eds:T Fenchel & K Sand Jensen), Naturen i Danmark. Gyldendal RN Glud (2006) Oxygen exchange; a key measure in marine biology requesting complementary technique EU, News letter RN Glud (2006) COBO modules used in collaboration with JAMSTEC, Japan: Towards biogeochemical observatories for benthic research EU, News letter RN Glud, M Middelboe (2006): Havbunden; et dynamisk levested for havbundens centrale aktører. Naturens Verden 11/12:2-10. PB Christensen, RN Glud, S Rysgaard (2008): Havisen i Arktisk: En vigtig brik i det globale klima. Naturens Verden 5:2-9. Rysgaard S, Sejr MK, Christensen PB, Born EW, Glud RN (2009): Det marine Miljoe i Naturen og klimaforandringerne i Nordoestgroenland (Eds: MC Forekammer, H Meltoft, M rasch). Aarhus Universitetsforlag National Television - DK RN Glud (1999) Interview in televison DR1 & DR2 Isgrænsen fantastiske liv Viden Om National Radio -DK S Rysgaard, RN Glud (2001) DR-P1 30 min udsendelse Bakteriernes liv i havis 14

RN Glud (2003) DR-P1 35 min udsendelse Grønlands varmeste kilde 15 RN Glud & M Middelboe (2005) DR-P1 1 times udsendelse Havbundens virus i serien Videnskabens Verden. RN Glud (2007) DR -P1 1 times udsendelse CO2 i de Arktiske oceaners dyb i serien Videnskabens Verden. RN Glud (2007 27 Juli) 10 min interview i P1-morgen Udforskning af dybhavet