ЪНИЗО ВПУЩГ ПЕВВРЕЕ, 2004, 65, 2, 135151 НеД 612:8216; 575:599323591 fl 2004 Ъ Е И Е ЙВ ИОЗ З 630091 З, М, 11 e-mail:rezzhan@fennsuru К 26052003 Й, fl, - fl,,,,, З fl А fl -, fl, fl fl А (- ), А fl fl Й (social learning), fl- fl fl - fl, fl fl Й - fl, -,, (- ),,, - fl - Аfl - fl -, (А, 1913; Й, 1922; З, 1948, 1973; Ж, 1970; М, 1976; Krebs, Dacies, 1978) А fl - fl, fl fl fl О fl -, fl Social learning 27- (Reznikova, 2001) д fl А - fl fl, fl, - flfl - fl Е Ч Й-б XIX, fl:,,, -,, Ч fl fl д fl,, fl,, -, fl -, (,, -,, ) - А fl - fl fl - В - fl В fl, - flfl,,, - З fl fl, flfl,, flfl 135
136 И flfl е, fl fl fl, fl fl: fl - fl, fl, - fl а - Е fl fl, Й fl -, fl fl flfl еfl fl -, fl - Е fl - fl fl -, fl Й, - fl flfl - - fl, fl fl fl (Х, 1977; К, 1983; а, 1991; Е, 1997; Й, Х, 1989; В, Й, 1998; И, 1998; У-, 2000; П, Й, 2001) Й, бfl fl Social learning -, - fl, -,, - fl (Heyes, 1994) Ч - fl flfl (imitation learning), - (observational learning) К - К flfl, -, fl,, fl- fl Ж - fl fl fl, fl flfl ( contagion - ) - fl social faclitation, social release (Zajonc, 1965; Р, 1983) А, flfl (, -, ) fl (), К fl fl -, К- fl,,,, ( ) - -, fl,, В - fl fl (stimulus enhancement): - - - И fl ( ) - - а fl (stimulus faclitation): fl fl - fl fl - (Spence, 1937; Thorpe, 1956) П - fl (emulation) еfl fl : flfl Ч б (Tomasello, 1990) fl fl -, fl fl fl, - fl А б - fl fl - З fl fl, -, fl Щ - fl,, -, fl З fl - fl -, fl fl М- fl fl fl - fl,, З fl - fl fl ( fl, Thorndike, 1911) Ч fl А
fl 137 fl flfl, - (Caldwell, Whitten, 2002) Й fl (Thorpe, 1956) З fl fl, - - - А fl fl (М, 1976; Р-, 1983; Ж, М, 1996), fl О -, fl fl А fl, fl, - fl fl,, fl, -,,, -, fl, fl-, Е, flfl - fl fl fl- Щ flfl fl- fl fl, fl fl, fl fl К fl (Hind, Fisher, 1951), (, 1961) - (Galef, 1992) Й - flflfl - (teaching, tutoring), fl, - Ч fl flfl : О А, О fl - А, fl fl К О,, - А, flfl, fl - fl, fl flfl, fl fl А - А fl () fl,, (Caro, Hauser, 1992) О - flflfl fl fl - fl flfl З fl - fl, fl,, - Мfl fl А fl -, fl fl, - fl З, - fl, - А, fl - fl А -, fl fl В-,, fl fl fl, - fl, (Oldfild-Box, 1970) А fl - fl а fl, fl - В,, fl, 15 Е fl flfl : -,, -, (М, М, 1982) Ж, flfl -, fl, -, -,, - fl, (Valsecehi et al, 1989) П, - fl fl (- ), - fl Й fl,, - (Ж, М, 1996) Й -, - fl, (Laland, Plotkin, 1992) Ж fl, - fl (Yeyes, 1994) Ж fl fl, fl, fl (Previde, Poli, 1996) А АБ Дfl (1940) fl
138 И fl 3040, fl, 23, fl- П, -, -, fl- (Darby, Riopell, 1959) О М (1972) -, fl - fl fl, fl fl К fl - fl fl, б, ОЗ К (1940), -, -, ; -,, fl, ; fl - fl fl О -, - fl (И, 1975, 1983; Reznikova J, 1982; Reznikova Zh, 2001) А, -, - fl - fl,, fl () Ч- fl - fl В, fl 10 - З, fl 2 А - fl, Щ - -,, flfl fl fl fl - а fl б, fl Ч fl Й -,,, К fl- fl, - fl, (), -,, fl К- flfl fl fl fl З, fl fl - fl, fl fl (Sherwin et al, 2002) А fl fl, б, - fl, -, - fl,, -,, fl (Suboski, Bartashunas, 1984) е,,, -, (Mason, 1988) еfl - - - В - fl, fl, fl Д-, - fl, (Galef, Giraldeau, 2001) А - - fl Н fl, fl, fl - Е fl fl, fl - (Rozin, Kalat, 1971; Д, 1989) З fl fl -, - а fl - Ч - Р (Galef, 1988), -- А - (, ), fl fl - fl fl Д - -
fl 139 -,, fl Ч flfl 12 В-, fl fl- fl, А - -, ( ) К fl fl, - fl fl - fl В, fl, -, - fl flfl fl -,, К, -,, -, - В - fl fl fl З, fl -, fl, - fl -, fl (Й, Д, 1985) М fl fl -, fl А - fl - -, еfl flfl fl fl, -,, fl fl, fl - fl, fl, fl - fl З - fl - Лfl fl fl, - fl, fl, -, fl, fl,, fl fl Ч,, fl (Visalberghi, Addessi, 2000) Н fl fl дfl (, - ), - fl А - В fl, fl fl - - fl И- fl fl, - fl, fl fl fl, fl, fl (Sherwin et al, 2002) Д fl :, - Щ, - flflfl К - fl fl, Р (Galef, 1988) Д, fl,, -,, fl fl И - Кfl -,,, fl fl,, - fl -, fl, fl е- (, ) -,, - fl, - Д fl, fl - Ч, fl, fl fl (-, ) еfl - fl - (Noble et al, 1999; Sherwin et al, 2002) fl fl fl,
140 И fl А, -, fl flflfl, fl fl flfl fl б,, - - -, fl - (Ж, 1970) Е - fl fl Д fl fl, -, - :,, fl, flfl В fl, -, fl Иfl flflfl, - fl fl Ч fl- - fl, - flfl, - -,, - З fl -, flfl, fl - flfl (Mineka, Cook, 1988) В А -,, fl, -, fl В, fl, fl, flfl,, fl Ч - fl fl - fl А - fl fl - - fl fl Й - Й fl, fl : fl, fl fl В, - fl - Ч fl (Dugatkin, 1996) Й П fl, fl : - ( fl -, flfl ) -, - fl В, fl (424%),, Щ - 40%,, - fl, fl Д fl Й fl,,, - fl - - Д ( ) б (Thorndike, 1911) fl fl fl : fl fl (Visalberghi, Fragaszy, 1990), - fl (Byrne, Tomasello, 1995), fl (Tomasello, 1996+ В fl- КВ Д, (KФhler, 1925), б (THorpe, 1956) -,, - ( fl), - В, - fl fl,, -, К fl -,, fl fl, З,, fl - fl - fl fl -, - ( : а, 1999) б - - fl - З,, fl, - fl fl Ч fl fl fl А fl -, fl, - fl, - fl З fl -, -
fl 141 (two-ways action/one outcome) Ж - fl В - б (Thorndike, 1911), fl - К fl - fl ( - fl, fl ),,, fl - fl, - fl М fl - -,, -, fl - К Еfl fl, - - fl М -, - fl, З - fl - ( ) А б fl fl, fl З fl,, а fl- - fl З, -,, - fl- Ч fl fl б е М - Melopsittacus undulasus:, fl, fl - fl, fl, - (Dawson, Foss, 1965) Ч fl fl fl fl Ж flfl А - - - fl К, fl fl е fl,, (Zentall, 1996) е, fl,, -, fl - fl (Campbtl et al, 1998) А - fl -,, fl fl, fl (Bugnyar, Huber, 1997) А - - -, fl ( - ) Ч fl, - fl А () - (Voelkl, Huber, 2000) К - Corvus corax (Fritz, Kortschal, 1999), fl (Akins, Zentall, 1996), (Byrn, Tomasello, 1995; Campbell, Heyess, 2002) - е -, fl fl б, - А (Whiten, 1998) fl -, -, fl - T- е - fl fl,, -,, - - ; fl,, fl fl, - fl К -, -, - fl fl О-,, flflfl, fl,,, fl, Н Д (KФller, 1925), е - -, fl Й- flfl, - (Tomasello, 1996; Whiten, 1998) Ж fl, fl,, - fl Д -, fl -
142 И, fl (Custance et al, 1999) В fl -, fl - fl б, - - fl (Fawcett et al, 2002) А, - Й -, - fl (fl ) fl - (fl ) б -, бfl - fl,, -, З fl, - fl К - Й,, - А, - fl (Huber et al, 2001) Кfl fl, - fl fl - ; fl, flfl fl е В -, А, - Й- fl, fl,,, - - В - -, fl, - З fl, В А,, - fl, - fl В fl fl fl, fl, З,, З Е - - fl Е fl - fl fl fl, Еfl, fl,, fl - fl - fl А fl, fl fl, А fl - - fl - - : fl fl, fl (fl- fl),,, fl И flfl А flfl, - (,, ) fl,, Ъ -, flfl,, Б flflfl (1970) fl, -, В, -,,, З - fl В,,, fl,,,,, fl б ПЕ Л (1947) - fl fl А - - Н, - fl, fl - fl fl,, - а fl - fl З fl - fl К
fl 143, fl, -, fl, -, fl, fl fl : fl, - (McGrew, 1992) А flfl fl flfl, - З fl, - fl, fl fl fl,, А fl,, fl, fl - fl б, 50-, fl, - fl fl fl -, fl (Church, 1957) А - -, fl а -, fl В- : К - ( fl ), Д fl, fl П fl, fl - И fl -, К, fl fl, fl (Brown, Laland,2002) Ч fl -, fl fl В fl, fl flfl е fl, fl fl б - fl flfl fl,, - fl (Cavalli-Sforza, Feldman, 1981; Boyd, Richerson, 1985) К - fl fl (Coussi-Korbel, Fragaszy, 1995) В,, fl fl fl fl : -,,, - К - :, fl- fl -, fl (Reader, Laland, 2000) Е fl - fl Steatocranus casuarius (Budaev et al, 1999), - (Benus et al, 1987), - (Verbeek et al, 1994), (Hessing et al, 1994) Н -, fl, - - (И, Иfl, 1990; И, З, 1998; Ryabko, Reznikova, 1996) еfl, - -, fl -, fl, fl, Е, fl fl - fl,, fl fl Н - - fl fl (Koolhaas et al, 1999) А fl - - fl (Pfeffer et al, 2002) Аfl, -, fl - fl Н, flfl - fl,, Ч, fl- -, fl- - а fl fl - А fl fl flfl fl
144 И (Lefebvre et al, 1997) О fl О () flflfl - А - fl -, fl fl (Caro, Hauser, 1992) Ж, flfl fl: 1) fl 2) - Кfl fl в fl, А fl fl fl - flflfl fl Аfl, fl,, fl А Д, flfl - fl,, - fl fl В, fl -, fl - Ч - fl fl - (Lopez G, Lopez D, 1985) (Ewer, 1969; З, 1998), - fl fl (Ewer, 1963; М, 1978) е - fl, fl fl fl - А fl,,, (И-, 2000, ) Л flflfl - Ч -Ч (Eibl-Fibesfeld, 1970) В, fl fl - fl,, fl fl fl, - fl Д fl fl З- fl fl fl,, fl fl - fl fl З,? З flflfl, -, fl? Ж, fl еfl fl- flfl fl fl, fl В flfl fl, fl, - еfl fl fl, fl, fl, fl - fl fl fl Й - : - fl fl, fl, fl- Е, fl, - fl - - fl (О, 1972; К, 1990) Щ- fl fl Д - Й (Caro, 1994) П fl, - fl fl 10 К, fl fl fl fl -, fl, - Йfl -, (Caro, 1980) В fl fl, fl, fl, fl, - И fl
fl 145 - А - fl, Ч fl - В - fl З fl, fl (б, 1970; Meinertzhagen, 1954; Beebe, 1960; McElroy, 1974, Newton, 1979) В fl,,, - fl fl И, - fl - fl fl - (Schaadt, Rymon, 1982) В fl fl -,, fl fl, -, fl - fl В flfl, Щ -, fl е - Й - fl fl - : fl (Fouts et al, 1982) А fl - 10 fl, - fl -, fl, fl (Boesch, 1991) К - К - 387 fl Ж - fl fl fl, fl б fl :, - fl, А - fl, (287 ) (316 ) З, - А - А, fl-,, fl fl, -, fl,,, fl А fl - - fl О (Midford et al, 2000) А - - В - : А fl, fl fl: fl - fl fl, fl fl, - Р fl Кfl fl - fl fl fl, flfl fl (Nagell et al, 1993): 1) fl - fl, fl fl; 2) fl- fl - fl- ; 3) fl ; 4) flfl е - fl fl -, fl, fl, е - А -, fl б - fl Л М (Fischer, Hind, 1949; Hind, Fisher, 1951), - fl, - fl fl fl - А - fl fl fl (Kawamura, 1963; Kawai, 1965) И - fl - Б fl -
146 И (De Waal, 1999) Кfl - fl б,, Д, fl fl, fl (Р-, 1990) Й, - fl fl, -, -, fl (Ж, 1986; Ж, М, 1997) А,, fl Д-, - fl fl - fl, fl, Е - fl,, - fl Ч - fl fl - fl, В,,,, fl - fl Ч fl, fl З, fl,,, (Zohar, Terkel, 1996) Ж - fl - fl - Й flflfl fl А fl, -, - Ж, fl, - - е fl - fl flfl -, б, Р (Galef, 1980), fl fl -, fl, - В, - fl fl -, fl И -, Р fl В flflfl -, fl flfl - fl, Ч - fl, В,, fl - fl, fl А (Sherry, Galef, 1984, 1990) -, fl fl fl fl fl fl flflfl fl- Щ,, - flfl, - fl А fl - Щ, А fl, Щ fl, fl - fl - А, fl, - fl В ( ) -,! Ч, -, - - А,,, fl М fl, fl flfl, flfl Кfl, - Ч - fl (а, 1986) А,, - Еfl,,, - - -
fl 147, Щ -, fl,, fl -, В - fl - fl, -,,,, - Д, fl, 7 fl О (McGrew, 1998, 2001; Whitten et al, 1999) е- ( ) 151 П 39 -, А, Ж - fl fl, Ч -, fl fl fl fl А д flflfl fl б, fl fl 1979 fl fl fl В-, flfl fl, - fl Йfl 20 80% fl Е- fl - - fl, fl, fl - Й fl flfl Ч fl,, fl-, fl - И - fl (Huffman, Hishie, 2001) З fl flfl, - -, - fl А - fl, flfl flfl, fl е flflfl - fl, fl -,, fl - В fl fl,, fl fl К,, fl, В fl А, О М (1977) fl - fl fl fl fl ( ),, В (fl), - а В fl, fl fl, -,, -, - - fl В fl - fl- fl Щ, Ч, fl, fl fl, - fl - е, fl - ( flfl ),, flfl fl- - З, flfl- fl К - : fl е -, fl - fl,,,, Чflfl fl, fl бfl fl - fl,, fl - fl fl Е- fl,, б в, -
148 И fl, flflfl, fl - В,, - fl,, -, - flfl flfl Д fl К, - - fl Нfl fl - fl fl, fl Й- (fl,, -, ), fl (, fl, - ) (, fl) fl -, fl - fl И ИММЕ ( 02-04- 48386) ЙКЕЙВД ЕбЩИОбНИ О е, 1972 Кfl Ж: Ж 215 П Ж, Й Щ, 2001 б - : fl - fl // Ъ 2001 T 62 1 C 7278 А А, 1913 П fl - ЙК ; Ж: Е- А б 2 435 c Р ЗЗ, 1990 вfl Д // А Д б 10 C 112115 Р ЖЩ, 1983 Й - fl : fl fl // Е Ж: АЕЗЕбЕ б 12 C 71150 Р е, 1992 Х : Ж: Ж 670 c а ОО, 1991 Вfl Ж: З 277 а ао, еfl ЖО, Ж ЗА, 1986 Й fl- // а б 65 10 C 15521559 а ао, К ЕЕ, И ЪЕ, 1999 В fl Ж: Е- ЖРН 383 c Е АА, 1997 А fl :, fl, fl Ж: Е- ДЖД Scientific Press 147 c Д ЩА, Ж ЗЗ, Х ЖЕ, 1989 В Ж: З, 250 c Дfl АБ, 1940 Е- fl fl // П 10 C 365372 ЖЩ, 1961 Йfl // fl б 1 : Е- РН Й 311 Д, 1970 Д fl Й Ж: а 195 c Ж ПК, 1970 а - fl // П- fl Ж: З Й 1236 Ж ЕИ, 1986 З - fl // IV А - б 3 Ж: З б 3 C 242244 Ж ЗЗ, М ЩЦ, 1996 В- -fl fl, Ж: О 225 c З ЙА, 1998 Аfl - Felis lynx, Felidae, Carnivora // а б 77 11 C 13171320 З ЗК, 1948 Чfl Ж: А 618 З ЗК, 1973 Й ( ) fl fl // Ъ - б 34 6 C 808817 В ВА, Й ЖО, 1998 Жfl - fl - fl Clethrionomys // К, - fl fl : Й / К Й АЩ, И АА, Й- ЖО Ж: З Й 3265 К АЙ, 1990 П Ж: О- 215 c К ЩЗ, 1983 К - fl fl Ж: З 423 c К ОЗ, 1940 А fl // еоз ЙЙЙИ б 27 2 C 240244 И ЪЕ, 1975 З - fl, // а б 54 7 C 10201031 И ЪЕ, 1982 Ж fl З : З 208 И ЪЕ, 2000 Е fl Ъ в 1 Ж: З 280 c И ЪЕ, 2000 Ж fl Ч -
fl 149 в 2 Ж: З 208 И ЪЕ, Иfl ПБ, 1990 б-- fl // Ъ б 51 5 C 601609 И ЪЕ, З бо, 1998 Е- // Н - б 118 3 C 345356 И АА, 1998 И - fl fl (Mustela putorius L) // К, fl fl : Й / К Й- АЩ, И АА, Й ЖО Ж: З Й 119132 Й ОЗ, 1922 Чfl Ж: ВРЕа 54 c Й ЦЖ, Х ЙЕ, 1989 З fl (Mus musculus) (Apodemus sylvaticus) - // а б 68 6 C 99110 Й АЩ, Д ЩА, 1985 Йfl - fl - // И- fl fl Ж: З Й 617 б З, 1970 В,, Ж: Ж, 334 c М ДЧ, 1976 В Ж: Е- ЖРН 287 c М ДЧ, М РР, 1982 Д - fl // А ЖРН Й 14 2 C 5561 М О, 1972 Кfl : З 230 c М О, 1977 К - fl : З 161 c М ЕЖ, 1978 б : Е- РН 182 c Л ПЕ, 1947 Д fl // б - б 15 Ж: Ж Й 3544 Х ЕО, 1977 Ч- fl Ж: З 260 c У ЗО, 2000 З fl- // Н б 120 1 C 7387 Akins CK, Zentall TR, 1996 Imitative learning in male Japanese quail (Coturnix japonica) using the two-action method // J Comp Psychol V 110 3 P 316320 Beebe FL, 1960 The marine falcons of the nortwest Pacific Coast // Condor V 62 P 145189 Benus RF, Koolhaas JM, Van Oortmerssen GA, 1987 Individual differences in behavioural reactio to a changing environment in mice and rats // Behaviour V 100 P 105122 Boesch C, 1991 Teaching among wild chimpanzees // Animal Behaviour V 41 P 530532 Boyd R, Richerson PJ, 1985 Culture and the Evolutionary Process Chicago: Univ Chicago Press 560 p Brown C, Laland KN, 2002 Social learning of a novel qvoidance task in the guppy: conformity and social release // Animal Behaviour V 64 P 4147 Budaev SV, Zworykin DD, Mochek AD, 1999 Consistency of individual differences in behaviour of the lionheaded cichlid, Steatocranus casuarius // Behavioural Processes V 48 P 4955 Bugnyar T, Huber L, 1997 Push or pull: an experimental study on imitaiton in marmosets // Animal Behaviour V 54 P 817831 Byrne RW, Tomasello M, 1995 Do rats ape? // Animal Behaviour V 50 P 14171420 Caldwell CA, Whitten A, 2002 Evolutionary perspectives on imitaiton: is a comparative psychology of social learning possible? // Animal Cognition V 4 5 P 193208 Campbell FM, Heyes CM, 2002 Rats smell: odour-mediated local enhancement, in a vertical movement two-action test // Animal Behaviour V 63 P 10551063 Campbell FM, Heyes CM, Goldsmith A, 1998 A demonstration of simultaneous stimulus learning and response learning by observation in the European starling using a two-object/two-action method // Animal Behaviour V 58 P 151158 Caro TM, 1980 Predatory behaviour in domestic cat mothers // Behaviour V 74 P 128147 Caro TM, 1994 Cheetahs of the Serengeti Plains: group living in an asocial species Chicago: Chicag Univ Press 320 p Caro TM, Hauser MD, 1992 Is there teaching in non-human animals? // The Quaterly Review of Biology V 67 P 151174 Cavalli-Sforza L, Feldman MV, 181 Cultural Transmission and Evolution: A Quantitative Approach Princeton: Princeton Univ Press 220 p Church RM, 1957 The procedures for establishment of imitative behaviour // J Compar Psychol V 50 P 315322 Coussi-Korvel S, Fragaszy DM, 1995 On the relation between social dynamics and social learning // Animal Behaviour V 50 P 14411453 Custance DM, Whiten A, Fredman T, 1999 Social learning of artificial fruit processing in enculturated capuchin monkeys // J COmpar Psychol V 113 P 1323 Darby CL, Rioppel AJ, 1959 Observational learning in the rhesus monkey // J Compar Physiol and Psychol V 52 1 P 94102 Dawson BV, Foss BM, 1965 Observational learning in budgeigars // Animal Behaviour V 13 P 47074 De Waal FBM, 1999 Cultural Prymatology Comes on Ages // Nature V 399 P 635636 Dugatkin LA, 1996 Interface between culturally based preferences and genetic preferences: female mate choice in Poecilia reticulata // Proc Nat Acad Sci USA V 93 P 27702773 Eibl-Eibesfeldt I, 1970 Ethology The Biology of Behaviour BY: Holt, Rinehart and Winston Press 630 p Ewer RF, 1963 The behaviour of the meerkat (Suricata suricatta) Schreber // J Tierpsyclol V 20 P 570607 Ewer RF, 1969 The instinct to teach // Nature V 222 P 698672 Fawsett TW, Skinner AMJ, Golsmith AR, 2002 A test of initative learning in starlings using a two-action method with an enhanced ghost control // Animal Behaviour V 64 P 547556
150 И Fischer J, Hunde RA, 1949 The opening of milk bottles by birds // British Birds/ V 42 P 347357 Fouts RS, Hursch AD, Fouts DH, 1982 Cultural transmission of a human language in a chimpanzee mother-infant relationship // Psychobiological Perspectives Chid Nurturance Series V 3 / Eds Fitzgerald HE, Mullins JA, Gage P NY: Plenum P 159193 Fritz J, Kortschal K, 1999 Social learning in common ravens, Corvus corax // Animal Behaviour V 57 P 785793 Galef BG, Jr, 1980 Division for food analysis of a possible case of social learning in wild rat // Ibid V 94 P 416425 Galef BG, Jr, 1988 Imitation in animals: history, definition and interpretation of data from psychological laboratory // Comparative social learning / Eds Zentall T, Galef BG, Jr Hillsdale (New Jersey): Erlbaum P 328 Galef BG, Jr, 1992 The question of animal culture // Human Nature 3 P 157178 Galef BG, Jr, Giraldeau LA, 2001 Social influence on foraging in vertebrates: causal mechanisms and adaptive functions // ANimal Behaviour V 61 P 315 Hessing MJC, Hagels AM, Beek JAM, van, Wiepkema PR, Schouten WPG, 1994 Individual behavioral and physiological strategies in pigs // Physiol and Behavior V 55 P 3946 Heyes CM, 1994 Social learning in animals: categories and mechanisms // Biol Review V 69 P 200231 Hinde RA, Fischer J, 1951 Further observations on the opening of milk bottles by birds // British Birds V 44 P 393396 Huber L, Rechberger S, Taborsky M, 2001 Social learning affects object exploraiton and manipulation on keas, Nestor notabilis // Animal Behaviour V 62 P 945954 Huffman MA, Nushie H, 2001 Stone handling, a two decade old behavioural tradition in a Japanese monkey troop // Advances in Ethology V 36 P 3536 Kawamura S, 1963 The process of sub-culture propagation among Japanese Macaques // Primate Social Behaviour / Ed Southwick CH NY: Van Nostrand P 8290 Kawai M, 1965 Newly-acquired Pre-Cultural Behavior of the Natural Troop of Japanese Monkeys on Koshima Islet // Primates V 6 1 P 130 KФhler W, 1925 The Mentality of Apes L: Routlegde and Kegan Paul Reprint in NY: Liverigh 1976 350 p Koolhaas JM, Korte SM, De Boer SD, Van Der Begt BJ, Van Reenen CG, Hopster H, De Jong IC, Ruis MAW, Blokhius HJ, 1999 Copying styles in animals: current status in behaviour and stress-physiology // Neuroscience and Behavioural Reviews V 23 P 925935 Krebs JR, Davies NB (eds), 1978 Behavioural Ecology An Evolutionary Approach Oxford: Blackwell Scientific Publ 482 p Laland KN, Plotkin HC, 1992 Further experimental analythis of the social learning and transmission of foraging information amongst Norway rats // Behavioural Processes V 27 P 5364 Lefebvre L, Whittle P, Lascaris E, Finkelstein A, 1997 Feeding innovation and forebrain size in birds // Animal Behaviour V 53 P 549560 Polez GC, Lopez D, 1985 Killer whales (Orimus orca) of Patagonia, and their behavior of intentional stranding while hunting near shore // J Mammal V 66 P 181183 Mason RJ, 1988 Direct and observational learning by redwinged blackbirds (Agelaius phoeniceus): the importance of complex visual stimuli // Social Learning: Psychological and Biological Perspectives / Eds Zentall TR, Galef BG, Jr Hillsdale (New Jersey): Lawrence Erlbaum P 99115 McElroy H, 1974 Desert Hawking Tuscon: Cactus Press 216 p McGrew WC, 1992 Chimpanzee Material Culture: Implications for Human Evolution Cambridge: Cambridge Univ Press 277 p McGrew WC, 1998 Culture in non-human primates? // Ann Rev Anthropology V 34 P 97115 McGrew WC, 2001 The Other Faunivory Primate Insectivory and Early Human Diet // Meat Eating and Human Evolution / Eds Graig B, Stanford H, Bonn T oxford: Oxford Univ Press P 160177 Meinertzhagen R, 1954 The education of young ospreys // Ibid V 96 P 153155 Midford PE, Hailman JP, Woolfeden GE, 2000 Social learning of a novel foraging patch in familie of free-libing Florida scrub-jays // Animal Behaviour V 59 P 1199 1207 Mineka S, Cook M, 1988 Social learning and the acquisition of snake fear in monkeys // Comparative social Learning / Eds Zentrall T, Galef BG, Jr Hillsdale (New Jersey): Erlbaum P 5173 Nagell K, Olguin RS, Tomasello M, 1993 Process of social learning in the tool use of chimpanzees (Pan troglodytes) and human children (Homo sapiens) // J Compar Psychol V 107 P 174186 Newton I, 1979 Population Ecology of Raptors Vermillion (South Dakota): Buteo Books 399 p Noble J, Tuci E, Todd PM, 1999 Social learning and information sharing: an evolutionary simulation model of foraging in Norway rats // Proc Fifth European Conference on Artificial Life (ECAL'99) / Eds Floreano D, Mondada F, Nicoud, J-D Belrin: Springer-Verlag P 514523 Oldfild-Box H, 1970 Comments ot two prelimibary studies of observation learning in the rats // J Compar Psychol V 116 1 P 125128 Pfeffer K, Fritz J, Kotrschall K, 2002 Hormonal correlated of being an innovative greylag goose, Anser anser // Animal Behaviour V 63 P 687695 Previde EP, Poli MD, 1996 Social learning in the golden hamster // J Compar Psychol V 110 P 203208 Reader SM, Laland KN, 2000 Diffusion of foraging innovation in the guppy // Animal Behaviour V 60 P 175180 Reznikova J, 1982 Interspecific communication between ants // Behaviour V 80 P 8495 Reznikova Zh, 2001 Interspecific and intraspectific social learning in ants // Advances in Ethology V 36 P 108109 Rozin P, Kalat JW, 1971 Specific hungers and poisoning as poisoning as adaptive specializations of learning // Psychol Review V 78 P 459486 Ryabko BYa, Reznikova ZhI, 1996 Using Shannon entropy and Kolmogorov complexity to study the communicative system and cognitive capacities in ants // Complexity V 2 2 P 3742
fl 151 Schaadt CP, Rymon LM, 1984 Innate fishing behavior of ospreys // Raptor Res V 16 P 6162 Sherry DF, Galef BG, Jr, 1984 Cultural transmission without imitation: milk bottle opening by birds // Animal Behaviour V 32 P 937938 Sherry DF, galef BG, Jr 1990 Social learning without imitation: more about milk bottle opening by birds // Animal Behaviour V 40 P 987989 Sherwin CM, Heyes CM, Nicol CJ, 2002 Social learning influences the preference of domestic hend for novel food // ANimal Behaviour V 63 P 933942 Spence KW, 1937 Experimental studies of learning and higher mental processes in infra-human primates // Psychological Bull V 34 P 806850 Suboski MD, Bartashunas C, 1984 Mechanisms for social transmission of pecking preference to neonatal chicks // J Experim Psychol V 10 P 182194 Thorndike EL, 1911 Animal Intelligence NY: Macmillan 560 p Thorpe WH, 1956 Learning and Instinct in Animals Cambridge: Harvard Univ Press 494 p Tomasello M, 1990 Cultural transmission in the tool use and communicatory signalling of chimpanzees? // Language and Intelligence in Monkeys and Apes / Eds Parker ST, Gibson KR Cambridge: Cambridge Univ Press P 274311 Tomasello M, 1996 Do apes ape? // Social Learning in Animals: The Roots of Culture / Eds Heyes CM, Galef BG, Jr NY: Acad Press, Inc P 319346 Valsecehi P, Mainardi M, Mainardi D, 1989 On the role of the demonstrator of the solution of a problem in the house mouse // Ethology, Ecology and Evolution V 1 1 P 213216 Verbeek MEM, Drent PJ, Wiepkema PR, 1994 Consistent individual differences in early exploratory behaviour of male great tits // Animal Behaviour V 48 P 1113 1121 Visalberghi E, Fragaszy D, 1990 Do monkeys ape? // Language and Intelligence in Monkeys and Apes / Eds Parker ST, Gibson KR Cambridge: Cambridge Univ Press P 247273 Visalberghi E, Adessi E, 2000 Seeing group members eating a familiar food enhances the acceptance of novel foods in capuchin monkeys // Animal Behaviour V 60 P 6976 Voelki B, Huber L, 2000 True imitation in marmosets // Animal Behaviour V 60 P 195202 Whiten A, 1998 Imitation of the sequential structure of actions by chimpanzees (Pan ptroglodytes) // J Compar Psychol V 112 P 270281 Whitten A, Goodall J, McGreew WC, Nishida T, Reynolds V, Syglyama Y, Tutin CE, Wranghan RW, Boesch C, 1999 Culture in chimpanzees // Nature V 399 P 682685 Zajonc RB, 1965 Social facilitation // Science V 149 P 269274 Zentall TR, 1996 An analysis of imitatice learning in animals // Social Learning in Animals: The Roots of Culture / Eds Heyes CM, Galef BG, Jr San Diego: Acad Press P 221243 Zohar O, Terkel J, 1996 Social and environmental factors modulate the learning of pine-cone stripping techniques by bkack rats, Rattus rattus // Animal Behaviour 1996 V 51 P 611618 Social Learning in Animals: Comparative Analysis of Different Forms and Levels Zh I Reznikova Institute for Animal Systematics and Ecology of SD of RAS, Novosibirsk State University, 11 Frunze Str, 630091, Novosibirsk Russia e-mail: rezzhan@fennsuru Social learning as one of the key concepts fo cognitive ecology includes different forms of behavioural displays from relatively simple, such as social release and stimulus enhancement up to teaching and cultural transmission in animal communities Rapid developments of this fiels resulted in some contradictions in methods and terminology In this review different forms and levels of social learning are analyzed Ecological aspects of social learning are connected with diet shaping, fear of predatros and mate choicethe first aspect is the most studied but still discussible Social learning being an intricate component of feeding behaviour matches with innate behaviour, imprinting as well as early associative learning Investigation of congnitive aspects of social learning going back to Thorndikes crucial question Do apes ape? are now developing into series of questions uncluding even: Do ants ape? Elaboration fo universal methods of comparative studying of social learning such as artificial fruit and two ways / one outcome has essentially enlightened these questions and made comparative analysis possible Large continuum of siplays of cognitive skills in social learning has been revealed in non-primate species One of the most disctussible issues in the role of socieal learning in distribution of innovations Many authors have investigated this intriguing aspect of animal behaviour in fdifferent ways, such as long fiels obeservations as well as laboratory e xperiments based on artificial innovators that is speciment specially taught by experimentalist Many impressive results were obtained; in particular it turned out in contradiction with some mathematical models that individuals in groups are rather different in their psychophysiological predisposition to innovative behaviour Role of teaching in such different forms of behaviour as shaping of species-specific behavioural patterns and spread of innovations is considered Although the majority of aminals in wild populations are not teachers and pulils, some cultural aspects of behaviour were recently revealed, mostly in primates At the same time some classical results cocerning cultural transmission of new patterns (for example, bottle-opening in tits) were experimentally revised Many problems still remain unsolved, in particular, how spread of innovations may favour prosperity of populations; to what degree behavioural peculiarities of logal groups may be determined by processes of social learning; which internal and expernal facotrs and under what circumstances invest into social learning in natural environment