ЗЩГИВМЕ ЕВВРЕҐЩЙДЕЩ ЖЩЛОЗЕ Ж КИВЕ АВ ЗВРВ АЗЕЖОЗЕБ (ОЗОЕ ЕҐЩЙДЕГ ВП ВИ)

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1 ЪНИЗО АЙХЩГ ЗЩИАЗВГ ЩБ Щ ЗВЙ Е, 2003, 53, 2, Н Д ъ, ЗЩГИВМЕ ЕВВРЕҐЩЙДЕЩ ЖЩЛОЗЕ Ж КИВЕ АВ ЗВРВ АЗЕЖОЗЕБ (ОЗОЕ ЕҐЩЙДЕГ ВП ВИ) 2003 ИЕ Ж Е ИОВ, Ж, К К А ъ - К, -, - Й ъ, ъъ В ъ -, - З, ъ, ъ - - Д :,,,,,, ЧЧР, АК, -, 1 А Е - - ъ, - - К -, [80, 112, 136] Въ - [32, 108], -- ъъ, - А ъ ъ -, ЗЙ К - - К - ъ - --, ъ - () Е ъ () ъ, В [32], ъ ъ - Н З З [108] :, А - ОЗ [5, 6, 18] ЦП Р [6] - - :, - Въ, ЦП АБ И [10], ъ, - Е, ЦП АБ И,, - - И

2 134, ъ -, Ж -- Щ [89], - ЗЙ, ъ ъ З ъ- 30, -, - М, - ъ- ъ - -, (АК), ъ В ъ - АК ъ (- ) П АК ъ- 60, -,, - ъ АК - Еъ АК - -- ъ К Й - [145] ( ), Й- [132] ( ), А Р [71] (, - ), ОЖ Е [14] ( ), И Й [141] (- ) Й Л [74] (-, ) К ЧО ъ ъ - - [2, 30, 60, 75, 95, 104] А ъ ъ, АК ъ,,, ъ- -, К Й - [145] АК - ъ ъ, -, ъ- -, ъ А Р [71], И Й - [141], Й Л [74] А З В Ж [22] -, -, -, ъ ъ АК - ъ - ъ, АК ъ -, - - Ч - АК,, - - ъъ, - АК - -, И100 ъ [73] О ъ ъ АК (N100 И100) - [33], - АК - ъ, N100 [66] К Й Л [76],, - АК, ъ, - - ъъ - ъ, : ъъ ъ ъъ З - ъ - ъъ [93]: V2 V4 -, ъ- 60 К-,, -

3 ЗЩГИВМЕ ЕВВРЕҐЩЙДЕЩ ЖЩЛОЗЕ Ж КИВЕ АВ ЗВРВ АЗЕЖОЗЕ 135 ъ К, ъ- ъ- ъъ - ( V2 V4) [38, 93, 101, 102, 113] А - ( V1), - ъ, ъ ъ- [73, 93, 94, 101, 113] А ъ ъ - - ъъ - (low-level), -, ъ - [94] К Д [54], ъ (top-down) ъ -,, ъ А Ж Дъ - [53] ъ, ъ, - : - V5,, ИВ (- ъ) А, - - ъ,, ъ, - -, - ъ ъ ъ П Ж [102], V1 ъ, ъ, - Р Х [138] -, - Е - - [144, 157] - V1 ( ъ,, - ) О Ж [97] - АК ъ - V1 P1 N1 - ъ -, В - АК, V1 - З П Ж Ц [41], ъ -,, - А ъ- ъ - ъ- А ъ И [127], ъ ( ъ ) ъ V1 - Й,, ъ - Й [30, 31, 125], ъ,, - - -, N100 АК [152] И -,, ъ - [66, 161] К ъ- И Д [81] -- ъ - - ъ А ъ Л Е-П Ж [79], -

4 ,, - ъ ъ -, ъ ъ- М - - ъ, ъ, - [105, 151], - ъ - Ч, ъ- ъ, ъ,, ъ ъ, ъ, (topdown), ъ ъ - ъ Й - ъ Н И - Д [54] ъ,,, -,, - (sites of attentional modulation) - (source of attentional modulation) Й - - [1, 4, 19, 28] -, ъ ъ- А - ъ ъ - ъ Ч, -, - ъ -, - - ъ Ж, - А ( 1) -, - -, - (connections) - ъ - ъ А ъ ъ [29, 45, 50, 54, 106, 121, 139, 143, 156, 160] И ъ,, [3, 19, 67, 107, 123], - ъ ъ, - metacognition [63, 137], - ъ,, ъ, Въ ъ - ъ -, ъ ъ- [19, 123], ъ -, А - - ъ ЧО [8, 22, 114, 152] Д, ъ ъ [30, 49, 53, 81, 152] Д, ъ ъ ъ Ч ъ ъ (go / no go) ъ [53, 61, 126, 131], ъ - Stroop, ъ,, - [34, 48] Й -, ъ ъ ъ - А ъ,

5 ЗЩГИВМЕ ЕВВРЕҐЩЙДЕЩ ЖЩЛОЗЕ Ж КИВЕ АВ ЗВРВ АЗЕЖОЗЕ 137 ъ - ( ankers), П Д [49], ъ К ъ ъ, ъ - ъ ъ К, ъ ъ - ъ, [48] К ъ - А, ъ - ъ [56, 64, 88], - - ъ - ъ - ъ ъ Б М [59] З , ъ А ъ Ч Д [84] c - - ъ ъ, ъ П, ъ - - ъ, ъ - ( ) -, - ( - ) А ъ, - В, (- - ) ( - ъ ), Ж К [121, 122] -, - ъ- ъ, - З () () ъ- - : (posterior) - (anterior) К ъ, ъ ъ- Д - -, ъ, ъ - Въ,, К Ж К, (topdown) ъ- ъ - - А, ъ- Ж К (Stroop ), -,, ъ, - ъ - А -, ъ,, ъ- - ъъ -, [52, 99, 139] А, -, Д Къ [124] - К, ъ -?,?,?, ъ-, А -, Д Къ, - - ъ, - ъ Н -, - - Аъ -, ъ, ъ- ъ - [160] Ч

6 138, -, - -, Д [51] К - А ъ-,, -, ъ ъ, Ч ъ, ъ О - [3, 107, 124], ( ОЙ П [3] - ъ ) Въ - - Е, - [70, 72, 107, 155], - - К -, А З [107] ъ ъ Ч К РИ [70, 72],, ъ Ж ъ- ъ, ъ [3, 12, 107], ъ-, - А - [107, 134] - ъ Й ОЙ П [3] ъ -- - ( - ) -, Ч - -, И - -, ъ -, - ъ К, - [42], ъ- ъ- [77] ъ [115] Й ъ -, ъ - [78] П - ъ ъ, -, - -, - [150] Й,, -,, - - А ( 2) ъ -,,, ъъ -, ъ- А :, - ъ, ъ - -? З,, ъ АК ЧО Д [15] -, -, -- - ъ- К ЧО Д, Д

7 ЗЩГИВМЕ ЕВВРЕҐЩЙДЕЩ ЖЩЛОЗЕ Ж КИВЕ АВ ЗВРВ АЗЕЖОЗЕ 139 ъ, ъ ъ - ъ ъ- - - Й, Ґ Е [142, 143, 162, 163] И ъ -, ъ, - А - ъ - - [133] А - ъ (, --, ) Ч - -, ъ ъ-,, - - ( - ) ( ) К - -, - - [54, 120], ъ - ъ И ъ- - -, И А [158] ъ - -, - ъ - ъ О ъ ъ- ъ П [43] - ъ - ъ ъ - ъ,, ъ [153] Д - ъ - (ъ - - ) [159] Н - (attentional gate) ъ ъ - ъ [98] К М Д [57], ъ, - - П Д А И [58], ъ Д,, Е ъъ - -, --, ъ ъ Е -, - АК, -, (supression) - ъ (facilitation) [76] М -, - [100] Ч, - ъъ - -, Еъ Въ ъ АК [11, 140] З - О Мъ ЗА ъ- [29] -, ъ ъ

8 ъ - - ъ, - Щ Ґ Е Й [162], Ж А [154] ъ, ъ ЧО ( ) ъ - ъ - Й ъ-, - А Й Ґ Е ( ) ъ Ж - - ъ [13] ИО [12], -,, ъ - Й ИО ъ - - ъ ъ - Ч, -, ИО, ъ -, ъ ъ - И - - [91, 146] В, -, - И - [90, 146] Е ъ - -,, [9, 35, 90, 92, 109] A Е [62] - -, (single neuron doctrine),, ъ, Зъ, ъ ъ ( ъ binding) - ъ Лъъ (D Hebb, 1949, [62]),, ъ ъ В Лъъ - ъ: ъ ъ - ъ (superposition catastrophe) А ъ M Оъc (M Abeles, 1982, [62]) Д Жъ (C von der Malsburg, 1986, [62]),, ъ- ъ - А ъ - A Е c [62] - - -, - (3070 Р) - К, - - ъ, - ъ И - ъ - - ъ (binding) [83] ЗЗ ЙА О [9],, - ъ, К, -, ъ -, - ъ, - ЧО -, ( ) Щ Зъ [110] - - Е ъ,

9 ЗЩГИВМЕ ЕВВРЕҐЩЙДЕЩ ЖЩЛОЗЕ Ж КИВЕ АВ ЗВРВ АЗЕЖОЗЕ 141 ъ, - -, ъ - - ъ - К И И Й [128] - ъ - ъ- - О З [109] ъ [92] О АК , Ґ М Д М- [65], N100 ъ, - И [90], -, - ъ - ъ - -, ъ - - Й - ъ -, - (single cognitive state),, - ъ, -, -, -, - ъ -, - - Д, ъ - --,, - Ж ъ, - -, ъ, О - ъ () Зъ, ъ -, ъ ъ , ъ, ъ- -, [3, 55, 121] Ж ъ -, ъ - - [54, И М - А, -, - -, ъ -, ъ Й -, -, () - :, ъ М - - ъ, ъ А ъ, -, ъ ъ, ъ - - З - Ж П О Аъ [40] -, - -, - ъ- (1525 Р)

10 142 О- (idling) [111, 118, 148] Ж -, ъ -, З Д- Х [85] ъ ЧЧР - ъ,, ъ ъ - К --, ъ - ъ К, - ъ arousal, - А - Ж Щ К [103] ъ ъ-, - А ъ ъ Р М [117] - -ъ ъ ъ - - Х [135], ЧО - - (attention), ъ -, - ъ (intention) А,, - ъ - -, (attention intention) ъ - - А ъ - - Ч - ъ Р Н [156] А Д- [82], ЧО - -, : ( 1- - ) - ( 2- ), - - Й М Й [91] - -, ъ,, ъ -,, - ъ, [147] И [149] К- - - [24, 149] А 8 13 Р - - ъ -, - [7, 87, 116] Е -, -, ъ- ( ), -, -, ъ [69] А ъ ЧО -, - [35] О - -, Щ П [35, 36],, - -, Р,, А - ъ -, - Р - ъъ - - ъ А - - -, ъ - -, ъ-

11 ЗЩГИВМЕ ЕВВРЕҐЩЙДЕЩ ЖЩЛОЗЕ Ж КИВЕ АВ ЗВРВ АЗЕЖОЗЕ 143 ъ К ъ,, ъ ъ ъ, ъ ъ ъ- Е -, ъ ъъ - - ъ - ъ ъ ъ - Ж, ъ ъ -, ъ, - ъ, - Ч -, - И ъ - А - ъъ К -, ъ - -, ъ - В -, - ъ А ъ Е, ъ- А - ъ ъ З Н [27], Й З Н ъ, А ъ ЧО Д- [16, 17] ъ - З -, - ъ Еъ -, ъ-, ъ - АК П К Л П [119] Е ъ ъ, ъ - ъ- А - ъ -, ъ ъ К АК, -, - О - АК ъ - [14] И - ъ,, ъъ - А - - Е -, ЧЖ И [25], И З [104], ЗА Й ВК [26] ъ- ъ - В - А ( [23]) И З - В, -, ъ ъ ъ ъ М, -, И З - Зъ ъ- - К И [129, 130] К И - ъ-, В ъ, - () ( -

12 144 ъ) - - [130] К, ъ, ъ ъ, А - ъ - ъ ъ - -- ъ [129] К, - ъ - К - -, - - ъ (, ), ъ - А К И ъ - ъ: -, ъ - - О, -, - Д К И, Е - - ъ -, ъ - ъ, - К И, Ґ П [45] - ъ ъ ъ- К,, -, - -- (loops),, ъ А Е ъ [46] З [23, 39, 68] - ЧО ъ- -, -, ЧЧР - - Н ъ, ъ - ъ, ъ ъ ЧО - - [47] А Щ П Л Й [37] ъ, - - ( ъ ) -, К - -ъ - N100 О ЧО ъ- ъ ъ [36] А c ъ - ЧО, - О, ЧО - ъ - - ъ И - ъ ъ ЧО А - - ЧЧР ЗВ Ж [22] ъ,, ЧО ъ - А -, - - И -ъ -

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19 ЗЩГИВМЕ ЕВВРЕҐЩЙДЕЩ ЖЩЛОЗЕ Ж КИВЕ АВ ЗВРВ АЗЕЖОЗЕ 151 thalamic reticular nucleus in the rat // J Neurosci 1999 V 19 P Weinberger DR A connectionist approach to the prefrontal cortex // J Neuropsychiatry Clin Neurosci 1993 V 5 P Woldorff MG, Gallen CC, Hampson SA et al Modulation of early sensory processing in human auditory cortex during auditory selective attention // Proc Nat Acad Sci USA 1993 V 90 P Yingling CD, Skinner JE Regulation of unit activity in nucleus reticularis thalami by the mesencephalic reticular formation and the frontal granular cortex // EEG a Clin Neurophysiol 1975 V 39 P Yingling ChD, Skinner JE Gating of thalamic input to cerebral cortex by nucleus reticularis thalami // Attention, Voluntary Contraction and ERP: Progress in Clinical Neurophysioljgy V1 / Ed Desmedt JE Basel: Kargel, 1977 P 7096 Р ъ - Х - М Й - ; Ж - ; ИБ ; ИМ - Neurophysiological Mechanisms of Voluntary Attention: A Review RI Machinskaya Institute of Developmental Physiology, Russian Academy of Education, Moscow The review is focused on attention as behavior-controling process Neurophysiological, electrophysiological and neuropsychological studies of different brain structures during voluntary attention are analyzed These data that selective voluntary attention modulates activity of sensory specific cortical involved in relevant signal processing Fronto-thalamic system consisting of prefrontal correx and thalamic vediodorsal nuclei is shown to be main source of top-down selective modulation of voluntary attention The reciew proposed the hypothetical model of selective cortical modulation during attention based upon the available data and evidence of own electroencephalografic studies Key words: voluntary perception, top-down control, prefronal cortex, thalamus, brain electrical activity, EEG, alpha rhythm, EP, psychophysiology