A Comparison of Aquatic Invertebrate Assemblages Collected from the Green River in Dinosaur National Monument in 1962 and 2001

Transcription

1 A Comparison of Aquatic Invertebrate Assemblages Collected from the Green River in Dinosaur National Monument in 1962 and 2001 Final Report for United States Department of the Interior National Park Service Dinosaur National Monument 4545 East Highway 40 Dinosaur, Colorado Report Prepared by: Dr. Mark Vinson, Ph.D. & Ms. Erin Thompson National Aquatic Monitoring Center Department of Fisheries and Wildlife Utah State University Logan, Utah January 2002

2 i Foreword The work described in this report was conducted by personnel of the National Aquatic Monitoring Center, Utah State University, Logan, Utah. Mr. J. Matt Tagg aided in the identification of the aquatic invertebrates. Ms. Leslie Ogden provided computer assistance. Several people at Dinosaur National Monument helped us immensely with various project details. Steve Petersburg, Dana Dilsaver, and Dennis Ditmanson provided us with our research permit. Ann Elder helped with sample archiving. Christy Wright scheduled our trip and provided us with our river permit. We thank them all for all their help and good spirit. We also thank Mr. Walter Kittams (National Park Service, Regional Office, Omaha Nebraska) and Mr. Earl M. Semingsen (Superintendent, Dinosaur National Monument) for funding the study and the members of the 1962 University of Utah expedition: Dr. Angus M. Woodbury, Dr. Stephen Durrant, Mr. Delbert Argyle, Mr. Douglas Anderson, and Dr. Seville Flowers for their foresight to conduct the original study nearly 40 years ago. The concept and value of long-term ecological data is often bantered about, but its value is never more apparent then when we conduct studies like that presented here. Additional copies of this report are available on the internet at: cover photographs: Top left - Green River in Lodore Canyon, bottom - sunset near Big Island, top right - Homoeoneuria alleni [Ephemeroptera: Oligoneuriidae] adult and nymph. Scenery pictures by E. Thompson. Insect photograph by M. Vinson.

3 1 Summary of 1962 study Flaming Gorge Dam is located approximately 45 miles upstream from Dinosaur National Monument. The dam was completed in December Prior to the closure of Flaming Gorge Dam, the National Park Service initiated a study (National Park Contract No ) to assess and survey the Green River and its surrounding environment. This study is described in detail by Woodbury (1963). This study was led by Dr. Stephen Durrant from the University of Utah. The expedition party sampled locations along the Green River from the Gates of Lodore downstream to Split Mountain from 20 July to 6 August Samples were also collected on the Yampa River near Deer Lodge and Echo Park. Data were collected on aquatic plants, aquatic invertebrates, fishes, as well as riparian and plants and animals. The objective of their aquatic invertebrate sampling was to determine the species present in the Green River prior to the closure of Flaming Gorge Dam. A synopsis of their sampling methods from Woodbury (1963) is as follows: At each site, samples were collected from all habitat types present. Samples were collected using a hand screen and a ¼ inch minnow seine. Abundance data is presented in the report, but this data should be evaluated cautiously, as the samples were qualitative in nature. It seemed that the invertebrates were likely removed from the debris in the field and identified back at the University of Utah. Drs. George Edmunds, Jr. and Arden Gaufin of the University of Utah consulted in the collection and identification of the aquatic invertebrates. We transcribed the aquatic invertebrate data in Woodbury s (1963) into our computer system. Where taxonomic changes had occurred since 1963, we changed their classification (invertebrate genus or species names) to those currently valid. Study design Our objective was to replicate the 1962 expedition as closely as possible. We collected samples at the same locations (Table 1), at the same time of the year (Table 2), using similar methods as the 1962 study.

4 2 Sampling Locations Table 1. Location of aquatic invertebrate sampling sites. Sites DINO-01 to DINO-06 were sampled in 1962 and Sites DINO-07 and DINO-08 were sampled only in Site ID Location Latitude Longitude Elevation (m) DINO-01 Green River at Lodore Ranger Station DINO-02 Yampa River at confluence with Green River DINO-03 Green River upstream from the Yampa River DINO-04 Green River at Steamboat Rock DINO-05 Green River at Rainbow Park DINO-06 Green River at Split Mountain Campground DINO-07 Green River at Rippling Brook Campsite DINO-08 Green River at Big Island Campsite Methods used in 2001 Locations and physical habitat The latitude and longitude of each site was determined using hand held global positioning units. Elevation was determined using U.S. Geological Survey 7.5 min topographic maps. Water and air temperatures were determined at the time of sampling using thermometers. The conductivity of the water was determined using an Oakton TDS Testr 40 hand held meter and the ph was determined using an Oakton ph Testr 2. Aquatic invertebrates Qualitative samples The objective of qualitative invertebrate collections was to collect as many different kinds of invertebrates living at a site as possible. Qualitative collections of invertebrates were done at all sites. Samples were collected with a rectangular kicknet (457 x 229 mm) with a 500 micron mesh net and by hand picking invertebrates from woody debris and large boulders. All available habitat types (e.g., riffles, po ols, back waters) were sampled and all samples were composited to form a single sample from each site.

5 3 Quantitative samples The objective of quantitative invertebrate sampling was to collect invertebrates from a fixed sampling area and estimate their relative abundances. Quantitative samples were collected using a modified Surber sampler (0.093 m 2 ) with a 500 micron mesh net. Samples were collected by disturbing the area immediately upstream of the net with our hands and scrubbing individual substrate particles within the sampling area and allowing the invertebrates and detritus to wash downstream into the net. Eight samples were collected throughout a sampling reach and composited to make a single sample of approximately 0.7 m 2 (8 samples x m 2 ). Laboratory methods The general procedures followed for processing invertebrate samples were similar to those recommended by the United States Geological Survey (Cuffney et al. 1993) and are described in greater detail and rationalized in Vinson and Hawkins (1996). Both qualitat ive and quantitative samples were identified in their entirety, i.e., all of t he organisms collected were identified. Organisms were identified to the lowest taxonomic level possible based on organism maturity, except for Chironomidae and some non-insect groups, which were identified to sub-family or Class or Order. Insects were most frequently identified to genus or family for small, immature, or damaged specimens. Species level identifications were based on identification keys, distributions records, and the author s experience. The number of individual invertebrates collected and identified in each sample is shown in Table 2. All identified invertebrates in each sample were composited into a single museum-grade glass screw-top vial with a polypropylene lid and polypropylene liner. Sample labels were written with fade proof permanent black carbon ink on waterproof paper. Information on each label included the sampling location, sampling date and a unique catalog number. Vials were filled with 70% ethanol. All samples were retained in our collection and will be sent to Dinosaur National Monument upon request. These samples will also be made available to other researchers upon request and permission from the National Park Service. Data analysis and presentation The primary objective of this study was to compare the invertebrate assemblages collected in 1962 to those collected in Data are provided on the occurrence and relative abundance

6 4 of aquatic invertebrates collected at each site for qualitative (1962 and 2001) and quantitative (2001) samples as well as composite (quantitative and qualitative samples) samples (2001) and a composite of all invertebrates collected by each study for all sampling locations. We used the data collected in 1962 as the basis for all comparisons presented here. We feel that our sampling effort was as great or greater than that done in Thus, if we failed to find a taxon, we think this suggests that this taxon may be extirpated from this reach of the river. Comparisons of taxa collected at individual sites should be done by comparing their data to the data from our qualitative samples, as these were done using similar methods, and presumably similar sampling effort, as was done in Table 2. Sampling dates, habitats sampled, type of sample collected, and the number of invertebrates identified in each sample. Data from 1962 is from Woodbury (1963). Sample ID is the National Aquatic Monitoring Center s identification number. Sampling Habitat Sample Invertebrates Sample ID Station date sampled type Identified DINO-01 07/28/2001 Riffle Quantitative DINO-01 07/28/1962 All present Qualitative DINO-02 07/30/2001 All present Qualitative DINO-02 07/30/2001 Riffle Quantitative DINO-02 07/24/1962 All present Qualitative DINO-03 07/30/2001 All present Qualitative DINO-03 07/30/2001 Riffle Quantitative DINO-04 07/30/2001 All present Qualitative DINO-04 07/30/2001 Riffle Qualitative DINO-04 07/27/1962 All present Quantitative DINO-05 07/31/2001 All present Qualitative DINO-05 07/31/2001 Riffle Quantitative DINO-05 08/05/1962 All present Qualitative DINO-06 07/31/2001 All present Qualitative DINO-06 07/31/2001 Riffle Quantitative DINO-06 07/23/1962 All present Qualitative DINO-06 07/22/1962 All present Qualitative DINO-07 07/30/2001 Aerial & riparian Qualitative DINO-08 07/31/2001 Aerial & riparian Qualitative 48

7 5 Results & Discussion Physical habitat conditions There appeared to be no large obvious differences in aquatic invertebrate habitat conditions between 1962 and Water temperatures which have changed substantially upstream, were just a few degrees different. From the general site descriptions provided in Woodbury (1963) substrate and riparian conditions also appeared similar between study periods. However, the reader should see work by Dr. Jack Schmidt and Mr. Paul Grams from Utah State University who documented geomorphological changes in this river corridor since the closure of Flaming Gorge Dam (e.g., Grams 1997). Table 1. Water temperatures, conductivity, and ph at locations sampled for aquatic invertebrates in 1962 and Data from 1962 is from Woodbury (1963). Air Water Conductivity Station Date Time (h) Temperature (C) µs cm -1 ph DINO-01 07/28/1962 **** ******************no data********************** DINO-01 07/28/ DINO-02 07/30/ DINO-03 07/30/ DINO-04 07/27/ DINO-04 07/30/ DINO-05 08/05/1962 **** ******************no data********************** DINO-05 07/31/ DINO-06 07/22/ DINO-06 07/31/

8 6 Aquatic invertebrate assemblages The aquatic invertebrate assemblages collected nearly 40 years apart were more similar than we anticipated. In a similar study conducted upstream, Vinson (2001) found little similarity between the pre- and post Flaming Gorge Dam invertebrate assemblages. In the stream reach immediately downstream from Flaming Gorge Dam, the number of macroinvertebrate genera declined from > 70 to < 30 following dam closure. In this study the differences were far less. In 1962 a total of 31 unique aquatic invertebrates were collected from 8 orders and 24 families. At the genus level they collected 2 Coleoptera, 4 Diptera (excluding Chironomidae), 12 Ephemeroptera, 3 Hemiptera, 3 Odonata, 1 Plecoptera, and 4 genera of Trichoptera. They did not report collecting any non-insect invertebrates. In 2001 we collected a total of 62 unique taxa from 8 orders and 31 families. At the genus level we collected 3 Coleoptera, 3 Diptera (excluding Chironomidae), 12 Ephemeroptera, 1 Hemiptera, 1 Odonata, 2 Plecoptera, and 6 genera of Trichoptera. We also collected 2 genera of gastropod, the common Physella [Gastropoda: Physidae], and a rare limpet, Ferrissia rivularis [Gastropoda: Ancylidae]. Individual taxa lists for each sample, sampling location, and study are provided in Appendices 1-3. Taxa found in 1962, but not in 2001 There were several taxa collected in 1962 that we did not collect in These were Zaitzevia [Coleoptera: Elmidae], Gyrinus [Coleoptera: Gyrinidae], Palpomyia [Diptera: Ceratopogonidae], Ephemerella inermis [Ephemeroptera: Ephemerellidae], Heptagenia elegantula [Ephemeroptera: Heptageniidae], Isonychia campestris [Ephemeroptera: Isonychiidae], Lachlania powelli [Ephemeroptera: Oligoneuriidae], Pseudiron [Ephemeroptera: Psuedironidae], Notonecta, [Hemiptera: Notonectidae], Rhagovelia distincta [Hemiptera: Veliidae], Argia [Odonata: Coenagrionidae], Gomphus intricatus, Ophiogomphus severus [Odonata: Gomphidae], Claassenia sabulosa [Plecoptera: Perlidae], Cheumatopsyche [Trichoptera: Hydropsychidae], Nectopsyche [Trichoptera: Leptoceridae]. The few other minor differences between the taxa lists were primarily at the genus or species level, and are likely due to the maturity of the specimens collected in 1962 versus 2001, e.g., Simulium. For the taxa described above that were collected in 1962, but not in 2001, we offer the

9 7 following ideas. The Coleoptera; Zaitzevia, Gyrinus, and Hemiptera; Notonecta and Rhagovelia distincta were likely still present in the river, but we failed to collect them. These taxa are widespread in their distributions, but can be elusive to collect and they generally occupy more specialized slow water habitats that we did not sample as intensively as the main river channel. For Diptera, we did not collect Palpomyia, but we did collect 2 other Ceratopogonidae; Dasyhelea and Probezzia, and we also collected several other Ceratopogonidae individuals that were too immature to positively identify to genus. Therefore, Palpomyia may be present in the river. At the present, we do not know enough about its habitat requirements to suggest otherwise. Many of the immature Simuliidae that we collected were likely Simulium. We frequently collect Simulium, upstream and downstream from the monument. Three of the five mayflies that we failed to collect, Isonychia campestris, Lachlania powelli and Pseudiron may be extirpated from the river. We not find them in this study and we have not collected them in over 8 years of sampling both upstream and downstream from Dinosaur National Monument. The other 2 mayflies, E. inermis and Heptagenia sp., are frequently collected by us both upstream and downstream from the monument, and we suspect they are still present in this reach of the river, but for some unexplainable reason we failed to collect them. The Odonates, Argia, Gomphus intricatus, and Ophiogomphus severus, are likely still present in the river as well. We have collected these genera upstream and downstream from Dinosaur National Monument over the last several years. The single stonefly collected in 1962, Claassenia sabulosa, was not found by us in this study. We have collected it in recent years in the Green River upstream from Flaming Gorge Dam, but not downstream from Flaming Gorge Dam, and it too may be extirpated from this reach of the river. One of the two caddisflies we did not collect, Cheumatopsyche, is commonly found by us upstream and downstream from Dinosaur National Monument. The other genera, Nectopsyche, has been collected by us upstream of Flaming Gorge Dam, near the town of Green River, Wyoming, during the last several years, but we have not found it downstream from the dam. It may be extirpated as well. While there are several taxa that may be extirpated from this reach of the river since the closure of Flaming Gorge Dam. The differences we observed over the last 40 years in this reach of the river are much less than those we documented in upstream reaches. The reason for this is

10 8 most likely because of the influence the unregulated Yampa River and other smaller tributary streams impart on the Green River downstream from their confluences. Opportunities for future work We feel that the work presented here is preliminary, but provides a solid foundation on which we can expand our understanding of the effects of the operation of Flaming Gorge Dam on the downstream Green River ecosystem. We demonstrated that the effects of Flaming Gorge Dam on downstream biota are certainly ameliorated in downstream reaches, but impacts of the dam are still discernable. These and additional data will be valuable to assess longer term impacts of the dam, especially as discharge and water temperature releases from the dam are altered in the future. In 2001, our laboratory had a fairly aggressive, but primarily unfunded, research program on the Green River. We collected aquatic invertebrates from near Fontenelle Dam downstream to the mouth of the Green River where it joins the Colorado River. In 2002 and future years we would like to acquire funding to expand our sampling on the Green River. We encourage the National Park Service to partner with us in this project.

11 9 Literature Cited Cuffney, T.F., Gurtz, M.E. and Meador, M.R Methods for collecting benthic invertebrate samples as part of the National Water-Quality Assessment Program. United States Geological Survey Open-File Report Grams, P. E Geomorphology of the Green River in Dinosaur National Monument. Unpublished Master s Thesis, Utah State University, Logan, Utah, USA. Vinson, M.R A history of aquatic macroinvertebrate assemblage changes downstream from a large dam. Ecological Applications 11: Vinson M.R. and C.P. Hawkins Effects of sampling area and subsampling procedures on comparisons of taxa richness among streams. The Journal of the North American Benthological Society 15: Woodbury, A. M., editor Studies of the biota in Dinosaur National Monument-Utah and Colorado. University of Utah, Division Biological Science, Institute of Environmental Biological Research, Miscellaneous Papers 1, Salt Lake City, Utah, USA.

12 Appendix 1. List of taxa collected in Dinosaur National Monument in Sampling locations are provided in Table 1. Sampling dates are shown in Table 2. indicates taxa collected in 1962 but not Arthropoda Insecta Coleoptera Elmidae Zaitzevia Gyrinidae Gyrinus Diptera Ceratopogonidae Palpomyia Simuliidae Simulium Tipulidae Hexatoma Tipula Ephemeroptera Baetidae Acentrella turbidia Baetis Ephemerellidae Ephemerella inermis Heptageniidae Heptagenia elegantula Rhithrogena undulata Isonychiidae Isonychia campestris Leptohyphidae Tricorythodes minutus Leptophlebiidae Choroterpes albiannulata Traverella albertana Oligoneuriidae Lachlania powelli Polymitarcyidae Ephoron album Pseudironidae Pseudiron Hemiptera Gerridae Gerris remigis Notonectidae Notonecta Veliidae Rhagovelia distincta Megaloptera Corydalidae Corydalus cognata Odonata Coenagrionidae Argia Gomphidae Gomphus intricatus Ophiogomphus severus Plecoptera Perlidae Claassenia sabulosa Trichoptera Brachycentridae Brachycentrus Hydropsychidae Cheumatopsyche Hydropsyche Leptoceridae Nectopsyche

13 11 List of taxa collected from Dinosaur National Monument in Sampling locations are provided in Table 1. Sampling dates are shown in Table 2. Annelida Oligochaeta Arthropoda Arachnida Trombidiformes Crustacea Amphipoda Ostracoda Insecta Coleoptera Elmidae Dubiraphia Microcylloepus Optioservus Diptera Ceratopogonidae Dasyhelea Probezzia Chironomidae Chironominae Orthocladiinae Tanypodinae Empididae Hemerodromia Simuliidae Tipulidae Ephemeroptera Ameletidae Ameletus Baetidae Acentrella turbida Camelobaetidius Caenidae Brachycercus likely prudens Heptageniidae Leucrocuta pertersi Rhithrogena Leptohyphidae Asioplax Tricorythodes Leptophlebiidae Choroterpes albiannulata Traverella albertana Oligoneuriidae Homoeoneuria alleni Polymitarcyidae Ephoron album Hemiptera Corixidae Gerridae Gerris Lepidoptera Pyralidae Petrophila Megaloptera Corydalidae Corydalus cognata Odonata Coenagrionidae Gomphidae Plecoptera Nemouridae Zapada Perlodidae Isogenoides Trichoptera Brachycentridae Brachycentrus Glossosomatidae Helicopsychidae Helicopsyche Hydropsychidae Hydropsyche Hydroptilidae Hydroptila Leptoceridae Oecetis Rhyacophilidae Rhyacophila Mollusca Gastropoda Basommatophora Ancylidae Ferrissia rivularis Physidae Physella Nematoda

14 Appendix 2. Taxa lists and relative abundances for each individual sample collected in 1962 and samples Green River at Lodore Ranger Station Taxonomic list and abundances of aquatic invertebrates collected 07/28/1962 at station DINO-01, Green River at Lodore Ranger Station. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Diptera Ceratopogonidae Palpomyia L 1 Ephemeroptera Baetidae Baetis L 9 Ephemeroptera Heptageniidae Heptagenia elegantula L 8 Ephemeroptera Leptohyphidae Tricorythodes minutus L 4 Ephemeroptera Leptophlebiidae Traverella albertana L 19 Ephemeroptera Oligoneuriidae Lachlania saskatchewanesis L 63 Ephemeroptera Polymitarcyidae Ephoron album L 1 Odonata Coenagrionidae Argia L 1 Odonata Gomphidae Stylurus intricatus L 14 Trichoptera Brachycentridae Brachycentrus L 3 Trichoptera Hydropsychidae Hydropsyche L 1 Trichoptera Leptoceridae Nectopsyche L 1 Total: 12 taxa Yampa River at confluence with Green River Taxonomic list and abundances of aquatic invertebrates collected 07/24/1962 at station DINO-02, Yampa River at confluence with Green River. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Diptera Ceratopogonidae Palpomyia L 1 Ephemeroptera Baetidae Acentrella turbidia L 1 Ephemeroptera Heptageniidae Heptagenia elegantula L 27 Ephemeroptera Heptageniidae Rhithrogena undulata L 4 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 28 Ephemeroptera Leptophlebiidae Traverella albertana L 58 Ephemeroptera Oligoneuriidae Lachlania saskatchewanesis L 2 Ephemeroptera Polymitarcyidae Ephoron album L 24 Ephemeroptera Pseudironidae Pseudiron A 1 Plecoptera Perlidae Claassenia sabulosa L 13 Trichoptera Brachycentridae Brachycentrus L 1 Trichoptera Hydropsychidae Hydropsyche A 1 Trichoptera Leptoceridae Nectopsyche L 2 Total: 13 taxa 163

15 Green River at Steamboat Rock Taxonomic list and abundances of aquatic invertebrates collected 07/27/1962 at station DINO-04, Green River at Streamboat Rock. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Diptera Ceratopogonidae Palpomyia L 1 Diptera Simuliidae Simulium L 4 Diptera Tipulidae Hexatoma L 1 Ephemeroptera Baetidae Baetis L 43 Ephemeroptera Heptageniidae Heptagenia elegantula L 38 Ephemeroptera Heptageniidae Rhithrogena undulata L 6 Ephemeroptera Isonychiidae Isonychia sicca campestris L 6 Ephemeroptera Leptohyphidae Tricorythodes minutus L 1 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 2 Ephemeroptera Leptophlebiidae Traverella albertana L 398 Ephemeroptera Oligoneuriidae Lachlania saskatchewanesis L 4 Ephemeroptera Polymitarcyidae Ephoron album L 126 Hemiptera Gerridae Gerris remigis A 3 Hemiptera Notonectidae Notonecta A 4 Hemiptera Veliidae Rhagovelia distincta A 31 Megaloptera Corydalidae Corydalus cognata L 5 Plecoptera Perlidae Claassenia sabulosa L 15 Trichoptera Hydropsychidae Cheumatopsyche L 1 Trichoptera Hydropsychidae Hydropsyche L 7 Trichoptera Hydropsychidae Hydropsyche A 3 Trichoptera Leptoceridae Nectopsyche L 2 Total: 20 taxa Green River at Rainbow Park Taxonomic list and abundances of aquatic invertebrates collected 08/05/1962 at station DINO-05, Green River at Rainbow Park. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Coleoptera Gyrinidae Gyrinus A 2 Diptera Tipulidae Tipula L 1 Ephemeroptera Baetidae Baetis L 38 Ephemeroptera Ephemerellidae Ephemerella inermis L 1 Ephemeroptera Heptageniidae Heptagenia elegantula L 40 Ephemeroptera Isonychiidae Isonychia sicca campestris L 5 Ephemeroptera Isonychiidae Isonychia sicca campestris A 15 Ephemeroptera Leptohyphidae Tricorythodes minutus L 6 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 14 Ephemeroptera Leptophlebiidae Traverella albertana L 5 Ephemeroptera Oligoneuriidae Lachlania saskatchewanesis L 18 Ephemeroptera Oligoneuriidae Lachlania saskatchewanesis A 6 Ephemeroptera Polymitarcyidae Ephoron album L 65 Odonata Gomphidae Ophiogomphus severus L 4 Odonata Gomphidae Stylurus intricatus L 2 Plecoptera Perlidae Claassenia sabulosa L 7 Trichoptera Hydropsychidae Hydropsyche L 1 Total: 15 taxa 230

16 Green River at Split Mountain Campground Taxonomic list and abundances of aquatic invertebrates collected 07/23/1962 at station DINO-06, Green River at Split Mountain Campground. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Coleoptera Elmidae Zaitzevia L 1 Diptera Ceratopogonidae Palpomyia L 1 Ephemeroptera Baetidae Baetis L 1 Ephemeroptera Heptageniidae Heptagenia elegantula L 15 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 5 Ephemeroptera Leptophlebiidae Traverella albertana L 11 Ephemeroptera Oligoneuriidae Lachlania saskatchewanesis L 1 Ephemeroptera Polymitarcyidae Ephoron album L 6 Odonata Gomphidae Stylurus intricatus L 2 Plecoptera Perlidae Claassenia sabulosa L 4 Total: 10 taxa Green River at Split Mountain Overflow Campground Taxonomic list and abundances of aquatic invertebrates collected 07/22/1962 at station DINO-06, Green River at Split Mountain Overflow Campground. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Coleoptera Elmidae Zaitzevia L 2 Ephemeroptera Baetidae Baetis L 18 Ephemeroptera Heptageniidae Heptagenia elegantula L 22 Ephemeroptera Isonychiidae Isonychia sicca campestris L 1 Ephemeroptera Leptohyphidae Tricorythodes minutus L 3 Ephemeroptera Leptophlebiidae Traverella albertana L 95 Ephemeroptera Oligoneuriidae Lachlania saskatchewanesis L 77 Ephemeroptera Polymitarcyidae Ephoron album L 96 Plecoptera Perlidae Claassenia sabulosa L 28 Trichoptera Brachycentridae Brachycentrus L 3 Trichoptera Hydropsychidae Hydropsyche L 26 Trichoptera Hydropsychidae Hydropsyche A 2 Trichoptera Leptoceridae Nectopsyche L 7 Total: 12 taxa 380

17 15 Data from 2001 samples Green River at Lodore Ranger Station Taxonomic list and abundances of aquatic invertebrates collected 07/28/2001 at station DINO-01, Green River at Lodore Ranger Station. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per meter squared. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. riffle Class: Arachnida Trombidiformes A 1 Class: Crustacea Amphipoda A 1 D Coleoptera Elmidae L 3 I,D Coleoptera Elmidae Optioservus L 3 Diptera P 1 D Diptera E 8 Diptera A 7 U,D Diptera Chironomidae P 1 Diptera Chironomidae Chironominae L 9 Diptera Chironomidae Orthocladiinae L 1 Diptera Chironomidae Tanypodinae L 1 Ephemeroptera L 5 D Ephemeroptera A 1 D Ephemeroptera Baetidae L 12 I,D Ephemeroptera Leptohyphidae Asioplax L 5 Ephemeroptera Leptohyphidae Tricorythodes L 4 Ephemeroptera Leptophlebiidae L 1 D Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 1 Hemiptera Corixidae E 1 Trichoptera Helicopsychidae Helicopsyche L 1 Trichoptera Hydropsychidae L 1 I Trichoptera Hydropsychidae Hydropsyche L 3 Trichoptera Hydroptilidae Hydroptila L 1 Phylum: Mollusca Class: Gastropoda Basommatophora Physidae Physella A 1 Total: 21 taxa 78

18 Yampa River at confluence with Green River Taxonomic list and abundances of aquatic invertebrates collected 07/30/2001 at station DINO-02, Yampa River at confluence with Green River. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Diptera Chironomidae Chironominae L 2 Diptera Chironomidae Tanypodinae L 2 Ephemeroptera Baetidae L 3 I Ephemeroptera Caenidae E 1 Ephemeroptera Heptageniidae Leucrocuta L 12 Ephemeroptera Leptohyphidae Asioplax L 1 Ephemeroptera Leptohyphidae Tricorythodes L 11 Ephemeroptera Leptophlebiidae L 2 I Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 34 R14 Ephemeroptera Polymitarcyidae Ephoron album L 4 R Odonata Gomphidae L 1 I Total: 11 taxa Taxonomic list and abundances of aquatic invertebrates collected 07/30/2001 at station DINO-02, Yampa River at confluence with Green River. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per meter squared. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. riffle Class: Crustacea Amphipoda A 1 D Ostracoda A 1 Coleoptera Elmidae Dubiraphia L 3 Diptera P 3 D Diptera A 1 U Diptera Ceratopogonidae L 8 I,U Diptera Chironomidae P 15 Diptera Chironomidae Chironominae L 78 Diptera Chironomidae Orthocladiinae L 4 Diptera Chironomidae Tanypodinae L 7 Ephemeroptera L 39 I,D Ephemeroptera Baetidae L 5 D Ephemeroptera Heptageniidae L 7 D Ephemeroptera Heptageniidae Leucrocuta L 9 Ephemeroptera Leptohyphidae Asioplax L 19 Ephemeroptera Leptohyphidae Tricorythodes L 79 Ephemeroptera Leptophlebiidae L 12 D Ephemeroptera Leptophlebiidae E 3 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 23 R6 Ephemeroptera Polymitarcyidae Ephoron album L 5 Hemiptera Corixidae L 9 I Hemiptera Corixidae A 1 U Hemiptera Corixidae E 1 Lepidoptera Pyralidae Petrophila L 1 Odonata Gomphidae L 1 I Trichoptera Hydroptilidae A 1 Trichoptera Leptoceridae Oecetis L 5 Total: 23 taxa 344

19 Green River upstream from the Yampa River Taxonomic list and abundances of aquatic invertebrates collected 07/30/2001 at station DINO-03, Green River upstream from the Yampa River. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Phylum: Annelida Class: Oligochaeta A 1 Class: Arachnida Trombidiformes A 1 Diptera Chironomidae Chironominae L 7 Diptera Chironomidae Tanypodinae L 15 Ephemeroptera Baetidae L 1 U Ephemeroptera Leptohyphidae Asioplax L 2 Ephemeroptera Leptohyphidae Tricorythodes L 11 Ephemeroptera Leptohyphidae Tricorythodes E 1 Ephemeroptera Leptophlebiidae L 1 D Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 4 Hemiptera Corixidae L 1 I Hemiptera Corixidae A 8 U Trichoptera P 3 Total: 11 taxa 56 17

20 18 Taxonomic list and abundances of aquatic invertebrates collected 07/30/2001 at station DINO-03, Green River upstream from the Yampa River. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per meter squared. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. riffle Phylum: Annelida Class: Oligochaeta A 17 Class: Arachnida Trombidiformes A 7 Coleoptera Elmidae L 1 I Coleoptera Elmidae Microcylloepus L 13 Coleoptera Elmidae Microcylloepus E 7 Coleoptera Elmidae Microcylloepus A 5 Coleoptera Elmidae Optioservus L 1 Diptera P 3 D Diptera A 1 U Diptera Chironomidae P 17 Diptera Chironomidae Chironominae L 294 Diptera Chironomidae Orthocladiinae L 28 Diptera Chironomidae Tanypodinae L 82 Ephemeroptera L 17 D Ephemeroptera E 7 Ephemeroptera Baetidae L 20 I,D,U Ephemeroptera Baetidae E 9 Ephemeroptera Heptageniidae L 8 I,D Ephemeroptera Heptageniidae Leucrocuta L 1 Ephemeroptera Leptohyphidae Asioplax L 125 Ephemeroptera Leptohyphidae L 12 I,D Ephemeroptera Leptohyphidae E 19 Ephemeroptera Leptohyphidae Tricorythodes L 75 Ephemeroptera Leptophlebiidae L 1 D Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 8 Ephemeroptera Leptophlebiidae Traverella albertana L 1 Hemiptera Corixidae L 3 I Hemiptera Corixidae E 1 Hemiptera Corixidae A 3 U Lepidoptera Pyralidae Petrophila L 1 Trichoptera P 12 Trichoptera L 1 D Trichoptera E 1 Trichoptera Brachycentridae Brachycentrus L 1 Trichoptera Hydropsychidae L 3 I Trichoptera Hydropsychidae Hydropsyche L 7 Trichoptera Hydroptilidae L 3 U Trichoptera Hydroptilidae A 3 Trichoptera Hydroptilidae Hydroptila L 7 Trichoptera Leptoceridae L 4 I Phylum: Nematoda A 8 Total: 30 taxa 841

21 Green River at Steamboat Rock Taxonomic list and abundances of aquatic invertebrates collected 07/30/2001 at station DINO-04, Green River at Streamboat Rock. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Diptera Chironomidae Chironominae L 4 Diptera Chironomidae Orthocladiinae L 2 Ephemeroptera Ameletidae Ameletus L 2 Ephemeroptera Baetidae L 13 I,U Ephemeroptera Heptageniidae L 2 I Ephemeroptera Heptageniidae Leucrocuta L 2 Ephemeroptera Heptageniidae Rhithrogena L 1 Ephemeroptera Leptohyphidae Asioplax L 2 Ephemeroptera Leptohyphidae Tricorythodes L 11 Ephemeroptera Leptophlebiidae L 1 I Ephemeroptera Leptophlebiidae E 2 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 38 Ephemeroptera Leptophlebiidae Traverella albertana L 10 R5 Ephemeroptera Polymitarcyidae Ephoron album L 5 Plecoptera Perlodidae L 3 I Trichoptera Glossosomatidae L 1 U Trichoptera Hydropsychidae Hydropsyche L 1 Trichoptera Leptoceridae L 1 I Total: 17 taxa

22 20 Taxonomic list and abundances of aquatic invertebrates collected 07/30/2001 at station DINO-04, Green River at Streamboat Rock. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per meter squared. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. riffle Class: Arachnida Trombidiformes A 8 Coleoptera Elmidae L 1 D Coleoptera Elmidae Microcylloepus L 1 Coleoptera Elmidae Microcylloepus E 4 Diptera P 1 D Diptera E 4 Diptera A 1 U Diptera Ceratopogonidae Dasyhelea L 1 Diptera Chironomidae P 4 Diptera Chironomidae Chironominae L 54 Diptera Chironomidae Orthocladiinae L 9 Diptera Chironomidae Tanypodinae L 4 Diptera Empididae Hemerodromia L 1 Diptera Simuliidae P 1 Diptera Tipulidae L 13 I,U Ephemeroptera L 11 D Ephemeroptera E 13 Ephemeroptera Ameletidae Ameletus L 4 Ephemeroptera Baetidae L 74 I,D Ephemeroptera Baetidae E 28 Ephemeroptera Baetidae Camelobaetidius L 51 R10 Ephemeroptera Baetidae Acentrella turbida L 17 Ephemeroptera Heptageniidae L 7 I,D Ephemeroptera Heptageniidae Leucrocuta L 5 Ephemeroptera Leptohyphidae Asioplax L 54 Ephemeroptera Leptohyphidae L 7 I,D Ephemeroptera Leptohyphidae E 12 Ephemeroptera Leptohyphidae Tricorythodes L 34 Ephemeroptera Leptophlebiidae L 3 D Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 12 Ephemeroptera Leptophlebiidae Traverella albertana L 9 Ephemeroptera Polymitarcyidae Ephoron album L 15 Hemiptera Corixidae E 1 Plecoptera Nemouridae Zapada L 1 Plecoptera Perlodidae L 3 I,U Plecoptera Perlodidae Isogenoides L 24 Trichoptera P 1 Trichoptera Glossosomatidae L 4 I Trichoptera Hydropsychidae L 4 I Trichoptera Hydropsychidae Hydropsyche L 3 Trichoptera Hydroptilidae L 1 U Trichoptera Hydroptilidae Hydroptila L 5 Trichoptera Leptoceridae L 8 I Trichoptera Rhyacophilidae Rhyacophila L 3 Phylum: Nematoda A 1 Total: 39 taxa 527

23 Green River at Rainbow Park Taxonomic list and abundances of aquatic invertebrates collected 07/31/2001 at station DINO-05, Green River at Rainbow Park. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Class: Arachnida Trombidiformes A 12 Coleoptera Elmidae L 1 I Diptera Chironomidae P 1 Diptera Chironomidae Chironominae L 8 Diptera Chironomidae Orthocladiinae L 1 Diptera Chironomidae Tanypodinae L 1 Ephemeroptera Baetidae L 1 U Ephemeroptera Baetidae Acentrella turbida L 3 Ephemeroptera Heptageniidae Leucrocuta L 1 Ephemeroptera Leptohyphidae Asioplax L 7 Ephemeroptera Leptohyphidae Tricorythodes L 27 Ephemeroptera Leptophlebiidae L 3 I Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 3 Ephemeroptera Leptophlebiidae Traverella albertana L 1 Ephemeroptera Polymitarcyidae Ephoron album L 10 Hemiptera Corixidae L 1 I Lepidoptera Pyralidae Petrophila L 2 Plecoptera Perlodidae L 1 I Trichoptera P 1 Trichoptera Brachycentridae Brachycentrus L 1 Trichoptera Hydropsychidae Hydropsyche L 2 Trichoptera Leptoceridae Oecetis L 1 Total: 22 taxa 89 21

24 22 Taxonomic list and abundances of aquatic invertebrates collected 07/31/2001 at station DINO-05, Green River at Rainbow Park. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per meter squared. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. riffle Phylum: Annelida Class: Oligochaeta A 1 Class: Arachnida Trombidiformes A 15 Coleoptera Elmidae L 1 I Coleoptera Elmidae Microcylloepus A 3 Coleoptera Elmidae Microcylloepus L 5 Diptera P 4 D Diptera E 11 Diptera A 7 U Diptera Ceratopogonidae L 1 U Diptera Ceratopogonidae Dasyhelea L 4 Diptera Ceratopogonidae Probezzia L 1 Diptera Chironomidae P 26 Diptera Chironomidae Chironominae L 267 Diptera Chironomidae Orthocladiinae L 27 Diptera Chironomidae Tanypodinae L 27 Diptera Tipulidae L 4 I Ephemeroptera L 9 I,D Ephemeroptera Baetidae L 47 I,D,U Ephemeroptera Baetidae E 7 Ephemeroptera Baetidae Acentrella turbida L 19 Ephemeroptera Caenidae E 4 Ephemeroptera Caenidae Brachycercus L 1 R Ephemeroptera Heptageniidae L 11 I,D Ephemeroptera Leptohyphidae Asioplax L 129 Ephemeroptera Leptohyphidae L 7 I Ephemeroptera Leptohyphidae E 16 Ephemeroptera Leptohyphidae Tricorythodes L 90 Ephemeroptera Leptophlebiidae L 7 D Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 11 Ephemeroptera Leptophlebiidae Traverella a;bertama L 28 Ephemeroptera Oligoneuriidae Homoeoneuria alleni L 1 Ephemeroptera Polymitarcyidae Ephoron album L 11 Lepidoptera Pyralidae Petrophila L 16 Odonata Gomphidae L 1 I Plecoptera Perlodidae L 1 I Trichoptera Brachycentridae Brachycentrus L 3 Trichoptera Glossosomatidae L 5 I,U Trichoptera Hydropsychidae L 4 I Trichoptera Hydropsychidae Hydropsyche L 12 Phylum: Nematoda A 3 Total: 35 taxa 848

25 Green River at Split Mountain Campground Taxonomic list and abundances of aquatic invertebrates collected 07/31/2001 at station DINO-06, Green River at Split Mountain Campground. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample of unknown area. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. multiple Coleoptera Elmidae Dubiraphia L 1 Coleoptera Elmidae Microcylloepus L 1 Diptera Chironomidae P 1 Diptera Chironomidae Chironominae L 12 Ephemeroptera Baetidae L 9 I,U Ephemeroptera Heptageniidae Leucrocuta L 1 Ephemeroptera Leptohyphidae Asioplax L 1 Ephemeroptera Leptohyphidae Tricorythodes L 12 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 25 Ephemeroptera Leptophlebiidae Traverella albertana L 1 Hemiptera Gerridae E 1 Phylum: Vertebrata Class: Osteichthyes L 1 I Total: 12 taxa Taxonomic list and abundances of aquatic invertebrates collected 07/31/2001 at station DINO-06, Green River at Split Mountain Campground. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per meter squared. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. riffle Phylum: Annelida Class: Oligochaeta A 4 Class: Arachnida Trombidiformes A 19 Coleoptera Elmidae L 7 I,D Coleoptera Elmidae E 3 Coleoptera Elmidae Microcylloepus L 17 Diptera P 1 U Diptera A 3 U Diptera Ceratopogonidae L 3 U Diptera Chironomidae P 12 Diptera Chironomidae Chironominae L 234 Diptera Chironomidae Orthocladiinae L 11 Diptera Chironomidae Tanypodinae L 12 Ephemeroptera L 16 I,D Ephemeroptera Baetidae L 20 I,D Ephemeroptera Baetidae E 5 Ephemeroptera Heptageniidae L 4 I Ephemeroptera Heptageniidae Leucrocuta L 1 Ephemeroptera Leptohyphidae Asioplax L 16 Ephemeroptera Leptohyphidae L 1 D Ephemeroptera Leptohyphidae E 12 Ephemeroptera Leptohyphidae Tricorythodes L 42 Ephemeroptera Leptophlebiidae L 4 D Ephemeroptera Leptophlebiidae E 3 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 17 Lepidoptera Pyralidae Petrophila L 7 Megaloptera Corydalidae Corydalus cognata L 1 Odonata Coenagrionidae L 3 I Trichoptera Hydropsychidae L 15 I Trichoptera Hydropsychidae Hydropsyche L 3 Phylum: Mollusca Class: Gastropoda Basommatophora Ancylidae Ferrissia rivularis A 1 Total: 25 taxa 497

26 Green River at Rippling Brook Campsite Taxonomic list and abundances of aquatic invertebrates collected 07/30/2001 at station DINO-07, Green River at Rippling Brook Campsite. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. Aerial & Life riparian Ephemeroptera Leptohyphidae A 26 Hemiptera Gerridae Gerris A 1 Total: 2 taxa Green River at Big Island Campsite Taxonomic list and abundances of aquatic invertebrates collected 07/31/2001 at station DINO-08, Green River at Big Island Campsite. Sample identification number is Life stage: L = larvae, P = pupae, A = adult, E = exuvia. NC = not calculated. Abundance data is presented as number per sample. Notes - identification to genus or species was not supported because: I - immature organisms, D - damaged organisms, M = poor slide mount, G = gender, U = indistinct characters or distribution, R = retained in our reference collection. Aerial & Life riparian Diptera A 8 U Ephemeroptera Leptohyphidae A 9 Ephemeroptera Leptophlebiidae A 40 Ephemeroptera Leptophlebiidae Choroterpes albiannulata L 3 Odonata Coenagrionidae A 1 U Trichoptera Hydropsychidae A 1 Trichoptera Hydroptilidae A 1 Total: 7 taxa 65

27 Appendix 3. Composite taxa list for each site sampled in Each taxa list was created by combining the data from 2 samples, a qualitative and a quant itative sample. Only a single sample was collected at DINO-01, DINO-07, DINO-08, so this data is not presented here, but is available in Appendix 2. List of taxa collected at station DINO-02, Yampa River at confluence with the Green River, on 30 July Arthropoda Crustacea Amphipoda Ostracoda Insecta Coleoptera Elmidae Dubiraphia Diptera Ceratopogonidae Chironomidae Chironominae Orthocladiinae Tanypodinae Ephemeroptera Baetidae Caenidae Heptageniidae Leucrocuta petersi Leptohyphidae Asioplax Tricorythodes Leptophlebiidae Choroterpes albiannulata Polymitarcyidae Ephoron album Hemiptera Corixidae Lepidoptera Pyralidae Petrophila Odonata Gomphidae Trichoptera Hydroptilidae Leptoceridae Oecetis

28 List of taxa collected at station DINO-03, Green River upstream from confluence with the Yampa River, on 30 July Annelida Oligochaeta Arthropoda Arachnida Trombidiformes Insecta Coleoptera Elmidae Microcylloepus Optioservus Diptera Chironomidae Chironominae Orthocladiinae Tanypodinae Ephemeroptera Baetidae Heptageniidae Leucrocuta petersi Leptohyphidae Asioplax Tricorythodes Leptophlebiidae Choroterpes albiannulata Traverella albertana Hemiptera Corixidae Lepidoptera Pyralidae Petrophila Trichoptera Brachycentridae Brachycentrus Hydropsychidae Hydropsyche Hydroptilidae Hydroptila Leptoceridae Nematoda

29 27 List of taxa collected at station DINO-04, Green River at Steamboat Rock, on 30 July Arthropoda Arachnida Trombidiformes Insecta Coleoptera Elmidae Microcylloepus Diptera Ceratopogonidae Dasyhelea Chironomidae Chironominae Orthocladiinae Tanypodinae Empididae Hemerodromia Simuliidae Tipulidae Ephemeroptera Ameletidae Ameletus Baetidae Acentrella turbida Camelobaetidius Heptageniidae Leucrocuta Rhithrogena Leptohyphidae Asioplax Tricorythodes Leptophlebiidae Choroterpes albiannulata Traverella albertana Polymitarcyidae Ephoron album Hemiptera Corixidae Plecoptera Nemouridae Zapada Perlodidae Isogenoides Trichoptera Glossosomatidae Hydropsychidae Hydropsyche Hydroptilidae Hydroptila Leptoceridae Rhyacophilidae Rhyacophila Nematoda

30 28 List of taxa collected at station DINO-05, Green River at Rainbow Park, on 31 July Annelida Oligochaeta Arthropoda Arachnida Trombidiformes Insecta Coleoptera Elmidae Microcylloepus Diptera Ceratopogonidae Dasyhelea Probezzia Chironomidae Chironominae Orthocladiinae Tanypodinae Tipulidae Ephemeroptera Baetidae Acentrella turbida Caenidae Brachycercus Heptageniidae Leucrocuta Leptohyphidae Asioplax Tricorythodes Leptophlebiidae Choroterpes albiannulata Traverella albertana Oligoneuriidae Homoeoneuria alleni Polymitarcyidae Ephoron album Hemiptera Corixidae Lepidoptera Pyralidae Petrophila Odonata Gomphidae Plecoptera Perlodidae Trichoptera Brachycentridae Brachycentrus Glossosomatidae Hydropsychidae Hydropsyche Leptoceridae Oecetis Nematoda