Braconidae (Hymenoptera) from Greece, 5.

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1 Notes fauniques de Gembloux, n 53 (2003) : Braconidae (Hymenoptera) from Greece, 5. by Jenő PAPP * Abstract A hundred forty-three braconid species, divided into eighteen subfamilies, are reported from Greece. Five species proved to be new to science: Triaspis graecus sp. nov., Opius (Adontopius) danielssoni sp. nov., Chelonus gozmanyi sp. nov., Ch. medinus sp. nov. and Ch. podlussanyi sp. nov. Eighty-one species are new to the fauna of Greece. With 67 original figures. Introduction A total of 411 braconid specimens served for the establishment of 143 species reported in the present account from the fauna of Greece. The species are subdivided into 18 braconid subfamilies. Five species are described as new to the science, their specification is presented in the abstract and, furthermore, they are related to their nearest allies. Eighty-one species proved to be new to the fauna of Greece. Vipio cinctellus Brullé, 1832 is redescribed and Opius (Opiothorax) graecus Papp, 1982 is revalidated. In the subsequent table, the braconid subfamilies are listed in alphabetical order completed with the species (the new ones in brackets) and the specimen numbers of the respective subfamilies dealt with here in the present paper : Subfamilies Species Specimens Agathidinae 9 28 Alysiinae Brachistinae 4 (1) 10 Braconinae Cardiochilinae 1 20 Cheloninae 39 (3) 147 Doryctinae 6 7 Euphorinae Gnamptodontinae 1 1 Helconinae 7 16 Homolobinae 1 1 Macrocentrinae 4 35 Meteorinae 7 20 Miracinae 1 1 Neoneurinae 2 4 Opiinae 5 (1) 12 Orgilinae 3 5 Rogadinae subfamilies 143 (5) species 411 specimens * Department of Zoology, Hungarian Natural History Museum, H-1431 Budapest, Pf. 137 (Hungary)

2 22 J. Papp The braconid wasps elaborated and reviewed here have come from and are deposited in nine institutions. Their abbreviated forms used in the faunistic list (after the number of the specimens) are presented in parenthesis as follows (see also Acknowledgement before Bibliography) : (in Berlin) = Museum für Naturkunde, Zologisches Museum, Berlin (in Budapest) = Hungarian Natural History Museum, Budapest (in Helsinki) = Department of Applied Zoology of the University, Helsinki (in Leiden) = Nationaal Natuurhistorisch Museum, Leiden (in London) = The Natural History Museum, London (in Lund) = Zoological Institute and Museum, Lund (in München) = Zoologische Staatssammlung, München (in Paris) = Museum National d Histoire Naturelle, Paris (in Wien) = Naturhistorisches Museum, 2. Zoologische Abteilung, Wien. Faunistic List In this list the subfamilies are ranged according to their systematic order, the genera and species alphabetically. Distributional and taxonomical remarks are added for those species to which it seems necessary. The added remarks originate from the respective publications cited in the Bibliography. DORYCTINAE Dendrosotinus titubatus Papp, (in Budapest): Epirus, Párga, 45 km NW Préveza, 9 23 June 1997, leg. R. Danielsson. Described from Greece, known from Italy and Israel. Heterospilus vilasi (Docavo, 1960) male new - 1 (in Budapest): Epirus, Párga, 45 km NW Préverza, 9-23 June 1997, leg. R. Danielsson. The male form is similar to the female. First flagellomere five times as long as broad. Vein 3-SR as long as 2-SR. First tergite 1.4 times as long as broad behind. Described and hitherto known from Spain. New to the fauna of Greece. Ontsira ignea (Ratzeburg, 1852) - 1 (in Budapest): Pentalofos, 18 Sept. 1997, leg. A. Podlussány. Distributed in the Palaearctic Region. New to the fauna of Greece. Nearest to Greece recorded from Croatia (Papp 1977: 106). Rhaconotus pictipennis (Reinhard, 1885) - 1 (in Lund): Epirus, Párga, 45 km NW Preveza, taken with yellow pan trap, 9-23 June 1997, leg. R. Danielsson. Up to now known from Germany and Hungary. New to the fauna of Greece. Rhoptrocentrus piceus Marshall, (in Budapest): Insel Andíparos (Kiklade), Olíaros, 2 Sept. 1925, leg. A. Schulz. Distributed in the Palaearctic Region, a less frequent species. New to the fauna of Greece; nearest to Greece recorded from Croatia (Papp 1977 : 107). Spathius rubidus (Rossi, 1794) - 1 (in Lund): Epirus, Párga, 45 km NW Préveza, 9 23 June 1997, leg. R. Danielsson. Frequent to common in the Holarctic Region. New to the fauna of Greece; nearest to Greece recorded from Croatia ("Lovrana" (Graeffe 1908: 143). GNAMPTODONTINAE Gnamptodon nieukerkeni Van Achterberg, (in Lund): Fokis, Giona Oros, 2 km N Sikea, 8 June 1982, leg. R. Danielsson. A very variable species. Body,

3 Notes fauniques de Gembloux, n 53 (2003) 23 antenna and legs black. Mesoscutum latero-posteriorly chagreened. Antenna with 19 antennomeres. Described from Greece (Parnassos Oros, 2 km W Summit, Voiotia) on the basis of by Van Achterberg (1983 : 34), reported from Poland (Tobias 1986 : 90). ROGADINAE Aleiodes (Neorhogas) ductor (Thunberg, 1822) - 1 (in München): Sykea, Sithonia, Chalkidike, 29 August 1977, leg. K. Warncke. Recorded from Greece (Papp 1985 : 218). Aleiodes (Aleiodes) gastritor (Thunberg 1822) - 2 (1 in Budapest, 1 in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Recorded from Greece (Papp 1985 : 218). Heterogamus testaceus Telenga, (in Lund): Pieria, Mt. Olympos, 4 km E Prioni, 22 June 1982, leg. R. Danielsson. 1 (in München) : Parnas, V. Delfi, 700 m, 28 May 1981, leg. Lukasch. Recorded from Greece (Papp 1985: 219). BRACONINAE Bracon (Glabrobracon) arcuatus Thomson, (in Budapest): Peloponesos, Prov. Messinia, Kardamili, 20 km SE Kalamaba, 20 May 1989, leg. M. J. Gijswijt. 1 (in Leiden): Ster. Ell., Etolia-Akar, Andiria, June 1989, leg. M. J. Gijswijt. Known in a few countries of Europe, nearest to Greece from Serbia and Croatia (Papp 1998/b: 148). New to the fauna of Greece. Bracon (Glabrobracon) cingulator Szépligeti, (in Lund): Epirus, Párga, 45 km NW Préveza, 9 23 June 1997, leg. R. Danielsson. Described from Russia ("Kasan"), known from Moldavia, Germany, Italy (Sardegna) and Korea. New to the fauna of Greece. Bracon (Glabrobracon) delibator Haliday, 1833 (=B. anthracinus Nees, 1834) - 1 (in Helsinki): Corfu, Acharavi, 24 April 1994, leg. M. Koponen. 2 (1 in Budapest, 1 in Helsinki): Corfu, Perama, 29 May 1978, leg. M. Koponen. 2 (in Leiden): Peloponesos, prov. Argolida, Arahneo, cca 500 m, 8 April 1989, leg. M. J. Gijswijt. Known from Greece (Papp 1990: 275). A common species in Europe. Its valid name "delibator" was recently notified (Van Achterberg 1997: 31). Bracon (Orthobracon) epitriptus Marshall, (in Helsinki): Corfu, Acharavi, 24 April 1994, leg. M. Koponen. Frequent to common in Europe. First reported from Greece by me (Papp 1999/b: 230). Bracon (Lucobracon) erraticus Wesmael, (in London): Ilia, Olympia, 4-11 July 1979, leg. M. C. Day. Known from Greece (Papp 1990: 275). Bracon (Lucobracon) fortipes Wesmael, (in London): Kifissia (Athenss), 8-25 June 1974, leg. L. A. Mound. In Europe a sporadic species, in the Balkan Peninsula fairly frequent. Bracon (Glabrobracon) fumatus Szépligeti, (in Leiden): prov. Attika, Kifissia (Athens), 4-5 April 1989, leg. M. J. Gijswijt. In Europe a sporadic species, nearest to Greece recorded from the former Yugoslavia (Papp 1998/b: 150). New to the fauna of Greece. Bracon (Glabrobracon) longulus Thomson, (4 + 1 in London, in Budapest): Mt. Pilion, Tsangarada, taken on Quercus sp., 20

4 24 J. Papp June Listed from seven countries in Europe: Sweden, Finland, Austria, Hungary, Italy, Croatia, Greece (Papp 1990: 276, 1998/b: 152, 1999/b: 232). Bracon (Glabrobracon) marshalli Szépligeti, (in Budapest): Ilia, Olympia, 4-11 July 1979, leg. M. C. Day. - Up to this time known only from Hungary. New to the fauna of Greece. Bracon (Glabrobracon) obscurator Nees, (in Leiden): Peloponesos, prov. Messinia, Proasto, 22 km SE Kalamata, 18 May 1989, leg. M. J. Gijswijt. A common European species. Recorded from Greece (Papp 1990: 277, 1999/b: 233). Bracon (Glabrobracon) parvulus Wesmael, 1838 (=B. fumipennis Thomson, 1892) - 1 (in Leiden): Peloponesos, prov. Korinthia, Stilia, 10 km SW Xilokostro, 5 June 1989, leg. M. J. Gijswijt. A fairly frequent species in Europe, nearest to Greece known from Macedonia (Papp 1998/b: 154). New to the fauna of Greece. Bracon (Bracon) pectoralis Wesmael, (in Helsinki): Corfu, Behitses, 1 : 3 June 1981, 1 : 24 May 1989, leg. M. Koponen. 1 (in Helsinki): Corfu, Perama, 30 May 1978, leg. H. et M. Koponen. Frequent to common in Europe, recorded from Greece (Papp 1990: 277, 1999/b: 233). Bracon (Pigeria) piger Wesmael, (in Helsinki): Corfu, Aghii Deka, 24 May 1981, leg. M. Koponen. 1 (in Helsinki): Corfu, Kanoni, 28 May 1978, leg. H. et M. Koponen. 1 (in Helsinki): Corfu, Perama, 28 May 1978, leg. H. et M. Koponen. A West-Palaearctic species, known from Greece too (Papp 1990: 277, 1999/b: 234). Bracon (Orthobracon) romani Fahringer, 1927 (=B. praetermissus auct. nec Marshall, 1885) - 1 (in Leiden): Peloponesos, prov. Arkadia, Ag. Petros, cca 1000 m, 24 April 1989, leg. M. J. Gijswijt. For a long decades a misunderstood species. Frequent to common in Europe; nearest to Greece known from Slovenia and Montenegro (Papp 1998/b: 155). New to the fauna of Greece. Bracon (Lucobracon) sphaerocephalus Szépligeti, (in Leiden): Peloponesos, "(Kornith.)", Tarsos, 17 May 1987, leg. H. Teunissen. 1 : Peloponesos, prov. Ilia, Nea Figalia, 28 May 1989, leg. M. J. Gijswijt. Recorded from Greece (Papp 1990: 278, 1999/b: 234). Bracon (Bracon) trucidator Marshall, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Distributed in Central and Souh Europe, recorded from Greece (Papp 1990: 278, 1999/b: 235). Bracon (Glabrobracon) variator Nees, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. A very common species in Europe, in Greece known from several localities. Bracon (Glabrobracon) variator var. bipartitus (Wesmael, 1838) (=var. rotundulus Szépligeti, 1901) - 2 (1 in Berlin, 1 in Budapest): Ost-Kreta, Kloster Ankárathos, Ep. Pedhiás, 28 May 1925, leg. A. Schulz. 2 (1 in Budapest, 1 in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Bracon (Glabrobracon) variator var. maculiger (Wesmael, 1838) - 1 (in Berlin): Rodi, Egeo, Mai 1939, leg. R. Meyer. 1 (in Leiden): Prov. Evrotonia, Kata fourka, 24 Sept. 1989, leg. M. J. Gijswijt. Bracon (Bracon) variegator Spinola, (in Leiden): Peloponesos, prov. Argolida, Arahneo, 500 m, 8 April 1989, leg. M. J. Gijswijt. A common species in Europe, known from Greece too.

5 Notes fauniques de Gembloux, n 53 (2003) 25 Coeloides sordidator (Ratzeburg, 1844) (=C. melanostigma Strand, 1918) - 1 (in Budapest): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. A frequent species in Europe. New to the fauna of Greece. Glyptomorpha pectoralis Brullé, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. In southern half of Europe a frequent and steppicolous species. Known from Greece (Papp 1990: 279, 1999/b: 236). Pseudovipio siculus (Marshall, 1888) - 1 (in Helsinki): Corfu, Aghii Deka, 24 May 1981, leg. M. Koponen. 1 (in Helsinki): Corfu, Perama, 3 June 1998, leg. M. Koponen. Recorded from Greece (Papp 1990: 180, 1999/b: 239). Vipio cinctellus Brullé, 1832: for its redescription see the chapter "Descriptions of the new species". Vipio humerator (A. Costa, 1884) (=V. frivaldszkyi Szépligeti, 1896) - 2 (1 in Budapest, 1 in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. A Ponto-Mediterranean species, recorded from Greece (Papp 1999/b: 239). Vipio longicauda (Boheman, 1853) (=Ichneumon nominator Fabricius, 1787) - 1 (in Lund): Fokis, Giona Oros, 2 km S Lefkadition, 8 June 1982, leg. R. Danielsson. 1 (in Lund): Pieria, Mt. Olympos, 4 km E Prioni, 22 June 1982, leg. R. Danielsson. A frequent species in the steppe and arboreal-steppe zone of the Palaearctic Region. Recorded from Greece under the name V. nominator (Papp 1990: 280). Vipio radiatulus Thomson, (1 in Budapest, 1 in Berlin): "Griechenland". 1 (in Berlin): Prov. Evrytania, M. Veluchi, 1700 m, 8 August, leg. Holtz. A rare species, known from Italy, Croatia (Istria) and Albania. New to the fauna of Greece. Vipio terrefactor (Villers, 1779) - 1 (in Helsinki): Corfu, Korisson, 4 June 1981, leg. H. et M. Koponen. 1 (in Helsinki): Lesbos, Ag. Isidoros, 17 May 1993, leg. H. Luoma. In the steppe and arboreal-steppe zone of the Palaearctic Region a less frequent to rare species. Nearest to Greece recorded from Croatia, Macedonia and Serbia (Papp 1998/b: 160). New to the fauna of Greece. HELCONINAE Aspicolpus carinator (Nees, 1814) (=Diospilus maximus Szépligeti, 1900) - 1 (in Budapest): Saloniki, 5 May 1932, leg. Paduschin Kattinger. A sporadic to frequent species in Europe. Nearest to Greece listed in Hungary (Papp 1998/a: 179). New to the fauna of Greece. Diospilus capito (Nees, 1834) (in Budapest): Crete, Heraklion, 5 May 1906, leg. L. Biró. 2 (1 in Budapest, 1 in Lund): Prov. Epidauros, Argolis, 17 May 1960, leg. M. Gyllenvärd. 1 (in Budapest): Prov. Lakonia, Monemvasia, 12 May 1979, leg. L. Gozmány. 1 (in Budapest): Prov. Lakonia, Sparta, Mt. Taygetos, 1000 m, 11 June 1980, leg. G. Christensen. In Europe a common species. Diospilus morosus Reinhard, (in Budapest): Marathon, 19 May 1977, leg. L. Zombori. Known from several countries in Europe, nearest to Greece from North Italy (Triest) (Graeffe 1908: 152). New to the fauna of Greece. Diospilus nigricornis (Wesmael, 1835) (=D. rufipes Reinhard, 1862) - 1 (in Budapest): Prov. Lakonia, Monemvasia, 10 May 1979, leg. L. Gozmány. Distributed

6 26 J. Papp in the Palaearctic Region, reported nearest to Greece from North Italy (Triest) and Slovenia (Tolmein) (Graeffe 1908: 152). New to the fauna of Greece. Diospilus oleraceus Haliday, 1833 (?=Bracon dispar Nees, 1812; = D. ruficornis Szépligeti, 1896) - 1 (in Lund): Nauplion, Argolis, 14 May 1960, leg. M. Gyllenvärd. In Europe frequent to common. Nearest to Greece recorded from Hungary (Papp 1998/a: 177). Diospilus productus Marshall, (in Budapest): Crete, Antr. Jovis, Mt. Ida, 1906, leg. L. Biró. 1 (in Budapest): Lake Volvi, 23 May 1977, leg. L. Zombori. 1 (in Budapest): Prov. Lakonia, Sparta, Mt. Taygetos, Anavriti, 1000 m, May 1979, leg. L. Gozmány. Hitherto reported from England and North Italy (Triest) (Graeffe 1908: 152). New to the fauna of Greece. Taphaeus hiator (Thunberg, 1822) - 1 (in Budapest): Crete, Heraklion, April 1906, leg. L. Biró. 1 (in Budapest): Sparta, Mt. Taygetos, 1400 m, May 1981, leg. A. Podlussány. Widely distributed in the Palaearctic Region, however, nowhere frequent. New to the fauna of Greece. BRACHISTINAE Schizoprymnus tantalus Papp, (5 in Lund and 2 in Budapest): Epirus Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Recorded in Turkey, Greece and Hungary (Papp 1998a: 169); in the Hungarian Natural History Museum there are specimens from Romania (Transylvania), Serbia, Macedonia, Italy and Algeria. It seems a frequent species in the southern half of Europe. Triaspis graecus sp. nov. - For its description see the chapter "Description of the new species". Triaspis luteipes (Thomson, 1874) - 1 (in Budapest): Epirus, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Known from Sweden, Italy and Switzerland. New to the fauna of Greece. Triaspis sulcatus (Szépligeti, 1901) - 1 (in Budapest): Viotia, Levadia, Tsoukalades, 5 June 1982, leg. R. Danielsson. Ovipositor sheath long, a bit longer than carapace; usually shorter. Hitherto known only in Hungary (Papp 1998a: 173). New to the fauna of Greece. METEORINAE Meteorus cinctellus (Spinola, 1808) - 1 (in Budapest): prov. Lakonia, waterfall between Nomia-Lyra, 20 June 1979, leg. L. Gozmány. Reported from eleven countries of Europe (Huddleston 1980: 25). New to the fauna of Greece. Meteorus gyrator (Thunberg, 1822) - 1 (in Budapest): prov. Lakonia, Mts Taygetos, 1000 m, 11 June 1980, leg. G. Christensen. 1 (in Budapest): prov. Lakonia, Monemvasia, taken with MV-lamp, 24 May 1979, leg. L. Gozmány et G. Christensen. Frequent to common in Europe. New to the fauna of Greece. Meteorus pulchricornis Wesmael, (in Budapest): prov. Lakonia, Monemvasia, taken with MV-lamp, 24 May 1979, leg. L. Gozmány et G. Christensen. Frequent in the western Palaearctic Region (Huddleston 1980: 45). New to the fauna of Greece. Meteorus rubens (Nees, 1812) - 3 (in Budapest): prov. Lakonia, 5 km S. of Monemvasia, 14 June 1980, leg. G. Christensen. 1 (in Budapest): prov. Thessalia,

7 Notes fauniques de Gembloux, n 53 (2003) 27 Kalambaka, Meteora, 9 September 1997, leg. A. Podlussány. Recorded from Greece (Papp 1990: 270). A common Meteorus species in the Palaearctic Region. Meteorus salicorniae Schmiedeknecht, (in Lund): Epirus. Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danilesson. Huddleston (1980: 48) reported it from Austria, Hungary,?Yugoslavia, and Bulgaria; Recorded from Greece (Papp 1990: 270). Meteorus tabidus (Wesmael, 1835) - 1 (in Budapest): prov. Halkidiki, Mt. Holomoudas, Taxiarhis, 22 May 1992, leg. I. Rozner. Reported from nine countries of Europe (Huddleston 1980: 50). New to the fauna of Greece. Meteorus versicolor (Wesmael, 1835) (in Budapest): prov. Lakonia, Monemvasia, taken with MV-lamp, 2 and 16 June 1979, leg. L. Gozmány et G. Christensen. 1 (in Budapest): prov. Lakonia, Mt. Taygetos, m, June 1979, leg. L. Gozmány et G. Christensen. 1 (in Budapest): prov. Lakonia, 5 km S of Monemvasia, 14 June 1980, G. Christensen. Recorded from Greece (Papp 1990: 270). EUPHORINAE Allurus lituratus Haliday, (in Budapest): Prov. Lakonia, Monemvasia, 17 April 1978, leg. J. Papp. 4 (in Budapest): environs of Stylis, 15 May 1977, leg. L. Zombori. Frequent to common in the western Palaearctic Region; new to the fauna of Greece. Allurus muricatus Haliday, (in Lund): Rhodes, Kattavia, Ag. Pavlos, 21 May 1983, leg. R. Danielsson. Recorded from Greece (Papp 1990: 270). Frequent to common in the western Palaearctic Region. Ancylocentrus edentatus (Haliday, 1835) - 2 (1 in Budapest, 1 in Lund): Rhodes, Kattavia, Ag. Pavlos, 21 May 1983, leg. R. Danielsson. A western Palaearctic species, in the Balkan Peninsula reported from Croatia (Istria; Graeffe 1908: 151) and Bulgaria (Zaykov 1979: 190). New to the fauna of Greece. Centites fuscipes (Nees, 1834) - 1 (in Lund): Rhodes, 3 km SE Kattavia, 21 May 1983, leg. R. Danielsson. A frequent species; reported from several countries of Europe, eastwards distributed as far as Georgia and Azerbaidzhan in the Caucasus Mts. New to the fauna of Greece. Leiophron deficiens (Ruthe, 1856) (1 and 1 in Budapest, 2 in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Widely distributed in the Palaearctic Region, albeit nowhere frequent. New to the fauna of Greece. Marshiella plumicornis (Ruthe, 1856) - 1 (in Budapest): Rhodes, Katavia, Ag. Pavlos, 21 May 1983, leg. R. Danielsson. Antenna with 17 antennomeres. The specimen represents an albanic form: head. mesosoma and petiole light brownish yellow, metasoma blackish brown; legs yellow; face and antennomeres lemon yellow, rest of antennomeres dark brown. Hitherto known sporadically in Germany, Hungary and West Kazakhstan. Microctonus aethiops (Nees, 1834) - 1 (in Lund): Fokis, Giona Oros, 3 k S Lefkadition, 7 June 1982, leg. R. Danielsson. 1 (in Budapest). Kavala, Pangeos Oros, 5 km NW Kipia, 20 June 1982, R. Danielsson. A Palaearctic species. New to the fauna of Greece.

8 28 J. Papp Microctonus lancearius Ruthe, (in Budapest): Fokis, Giona Oros, 3 km S. Lefkadition, 7 June 1982, leg. R. Danielsson. 1 (in Lund): Fthiotis, 3 km SW village Timfritos, 10 June 1982, leg. R. Danielsson. Described by Ruthe from Germany, reported from Macedonia and Serbia (Kosovo) (Papp 1973: 7). New to the fauna of Greece. Perilitus dubius (Wesmael, 1835) - 1 (in Lund): Fokis, 6 km SE Physkeis, 7 June 1982, leg. R. Danielsson. 1 (in Lund). Fthiotis, Iti Oros, 4 km NNE Kaloskopi, cca 1000 m, 9 June 1982, leg. R. Danielsson. Rather sporadic in Europe, eastwards distributed as far as the Mts Ural (Russia) and Kazakhstan. New to the fauna of Greece. Peristenus grandiceps (Thomson, 1891) - 1 (in Budapest): Crete, Canea, March 1906, leg. L. Biró. A western Palaearctic albeit not a frequent species. New to the fauna of Greece. Peristenus pallipes (Curtis, 1833) - 1 (in Lund): Trikala, Panagia, 15 June 1982, leg. R. Danielsson. One of the most frequent euphorine species in the Holarctics. In the Balkan Peninsula reported from Croatia (Istria; Graeffe 1908: 149) and Albania (Fahringer 1924: 104). New to the fauna of Greece. Pygostolus falcatus (Nees, 1834) - 1 (in Budapest): prov. Lakonia, Monemvasia, 10 May 1979, leg. L. Gozmány. 1 (in Budapest): prov. Lakonia, Mts Taygetos, m, taken with MV-lamp, June 1979, leg. L. Gozmány et G. Christensen. Frequent to common in Europe. New to the fauna of Greece. HOMOLOBINAE Homolobus (Apatia) truncator (Say, 1828) - 1 (in Budapest): Athens, Blaca, 18 October 1980, leg. L. Peregovits. Recorded from Greece (Papp 1985: 220). MACROCENTRINAE Macrocentrus (Macrocentrus) bicolor Curtis, (in Budapest): prov. Lakonia, Mts Taygetos, 1000 m, 11 June 1980, leg. G. Christensen. Farther three localities in Greece were presented by Haeselbarth (1978: 28 29). Distributed in the Palaearctic Region. Macrocentrus (Amicroplus) collaris (Spinola, 1808) - 1 (in Budapest): Alexandroupolis, 23 May 1977, leg. L. Zombori. 1 (in Budapest): Athens, Kifissia, 200 m, 1 April 1978, leg. J. Papp. 1 (in Lund): Iannina, 4 km N Metsovon, 16 June 1982, leg. R. Danielsson. 2 (in Lund): Crete, 1 km N Malia, taken at light, June 1979, leg. L. Cederholm (in Lund): Crete, 1 km N Malia, swept along beach, 30 June 1979, leg. L. Cederholm (in Budapest): prov. Lakonia, Monemvasia, taken with MV lamp, 1 April-16 October 1979, leg. L. Gozmány et G. Christensen. 1 (in Berlin): East Crete, Iraklion (Candia), 23 June 1925, leg. A. Schulz. A frequent to common species in Europe. Macrocentrus (Amicroplus) flavus S. van Vollenhoven, 1878 (=M. turanicus Telenga, 1950) - 1 (in Budapest): prov. Lakonia, Monemvasia, 15 September 1979, leg. G. Christensen. First reported from Greece (Crete) by van Achterberg (1993b: 39). Distributed sporadically in the western Palaearctic Region. Macrocentrus (Amicroplus) resinellae (Linnaeus, 1758) - 1 (in Budapest): prov. Lakonia, Mts Taygetos, m, taken with MV-lamp, June 1979, leg. L. Gozmány et G. Christensen. First reported from Greece by Achterberg (1993b: 55).

9 Notes fauniques de Gembloux, n 53 (2003) 29 CHELONINAE Ascogaster bicarinata (Herrich-Schaeffer, 1838) - 2 (in Budapest): Athens, Mt. Hymettos, Kaisarine, April 1931, leg. J. Fodor. 2 (in Budapest): Crete, Antr. Iovis, Mt. Ida, 1906, leg. L. Biró. 3 (1 in Budapest, 2 in Lund): Fokis, Giona Oros, 2 km S Lefkadition, 8 June 1982, leg. R. Danielsson. 1 (in Budapest): Kephallenia, leg. Paganetti. 1 (in Lund): Trikala, Orthovounion, 15June 1982, leg. R. Danielsson. 1 (in Lund): Trikala, Panagia, 15 June 1982, leg. R. Danielsson. First reported from Greece by Huddleston (1984: 354); a sporadic to frequent species in southern half of Europe. Ascogaster caucasica Kokujev, 1895 (=A. rostratus Szépligeti, 1896) - 1 (in Budapest): Prov. Attiki, Boulogméne, 13 April 1931, leg. J. Fodor (in Budapest): Prov. Attiki, Marathon, 21 April 1978, leg. J. Papp. 1 (in Budapest): Prov. Attiki, Paiania, 100 m, 3 April 1978, leg. J. Papp. 1 (in Budapest): Athens, Mt. Hymettos, Kaisarine, April 1931, leg. J. Fodor. 3 (in Budapest): Boiotia, Oinoi, 8 May 1981, leg. I. Rozner. 1 (in Lund): Fokis, Giona Oros, 2 km S Lefkadition, 8 June 1982, leg. R. Danielsson (in Budapest): prov. Thessaloniki, Asprovalta, 15 May 1995, leg. J. Papp. 1 (in Budapest): prov. Thessaloniki, Mikri Volvi, 21 May 1995, leg. A. Orosz. Huddleston (1984: 354) listed its five localities in Greece. In southern half of Europe widely distributed. Ascogaster quadridentata Wesmael, (in Lund): Viotia, Arachova, 6 June 1982, leg. R. Danielsson. Frequent to common in the Palaearctic Region. Ascogaster scabricula (Dahlbom, 1833) - 1 (in Budapest): Viotia, Arachova, 6 June 1982, leg. R. Danielsson. Reported from Belgium, France and Sweden (Huddleston 1984: 379). New to the fauna of Greece. Ascogaster varipes Wesmael, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. 1 (in Budapest): near Lithoron, Mts Olympos, 15 May 1977, leg. L. Zombori. Frequent to common in Europe. New to the fauna of Greece. Chelonus annulipes Wesmael, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Distributed in Europe. New to the fauna of Greece. Chelonus asiaticus Telenga, (in Budapest): Crete, Omalos, 16 July 1906, leg. L. Biró. 1 (in Lund): Evritania, Mt. Timfritos, 2 km N Karpinisi, 11 June 1982, leg. R. Danielsson. Distributed in Mongolia, Kazakhstan, Russia, Slovakia and Hungary (Papp 1996: 137). New to the fauna of Greece. Chelonus caradrinae Kokujev, (in München): N Ioannina, Papingon, 1000 m, 21 August 1977, leg. K. Warncke. Frequent in the western half of the former USSR, known from Hungary (Papp 1996: 138). New to the fauna of Greece. Chelonus carbonator Marshall, (1 in Budapest, 3 in Lund): Crete, Malia Beach near hotel, 12 May 1979, leg. R. Danielsson. 1 (in Lund): Crete, 1 km SE Molos, 4 km SW Malia, 600 m, swept in vegetation, 27 June 1979, leg. L. Cederholm. Frequent to common in Europe; its female is very rare. New to the fauna of Greece. Chelonus corvulus Marshall, (in Budapest): Crete, Antr. Iovis, Mt. Ida, 1906, leg. L. Biró. 1 (in München): Crete, Heraklion, 25 May 1963, leg. M. Schwarz. Frequent to common in Europe.

10 30 J. Papp Chelonus gozmanyi sp. nov.: For its description see the chapter "Description of the new species". Chelonus inanitus (Linnaeus, 1767) - 2 (in Budapest): Corfu, leg. Paganetti. 1 (in Budapest): Crete, prov. Irakleion, Amoudara, 9 May 1993, leg. A. Orosz. 1 (in Budapest): Prov. Drama, Mt. Falakova, 24 May 1995, leg. I. Rozner. 1 (in Budapest): prov. Thessaloniki, Skolarion, 21 May 1995, leg. A. Orosz. Widely distributed in the Palaearctic Region, nearest to Greece reported from Bulgaria (Zaykov 1978: 231). Chelonus medinus sp. nov.: For its description see the chapter "Description of the new species". Chelonus obscuratus Herrich-Schaeffer, (in Budapest): Corfu, Kombitsi, 27 June 1995, leg. Zs. Czetõ. 1 (in Budapest): Crete, Omalos, 18 August 1906, leg. L. Biró. 1 (in Budapest): Parnassos ("Ch. inanitus var. annulatus Nees" det. Szépligeti). 1 (in Budapest): prov. Thessaloniki, Mavrouda, 23 May 1995, leg. A. Orosz. The male specimen from Crete represting a melanic form: hind femur blackish and only apically reddish yellow, carapax entirely black (i.e. without a pair of basal yellow maculae). - Distributed in Europe, a sporadic to frequent species. New to the fauna of Greece. Chelonus oculator (Fabricius, 1775) (=Ichneumon oculator Panzer, 1779,?=Sigalphus mutabilis Nees, 1816) (In Budapest): Corfu, leg. Paganetti. 1 (in Budapest): Corfu, Kombitsi, 27 June 1995, leg. Zs. Czetõ. 1 (in Budapest): Crete, Antr. Iovis, Mt. Ida, 1906, leg. L. Biró. 1 (in Budapest): Crete, Heraklion, 5 May 1906, leg. L. Biró. 2 (in Lund): Crete, Malia Beach near hotel, 12 May 1979, leg. R. Danielsson. 2 (1 in Budapest, 1 in Lund): Crete, 1 km SE Molos, 4 km SW Malia, 600 m, swept in vegetation, 27 June 1979, leg. L. Cederholm. 1 (in Budapest): Kavaia, Keramoti, 20 June 1982, leg. R. Danielsson. 1 (in Lund): Rhodos, 4 km SW Lardos, 19 May 1983, leg. R. Danielsson. Frequent to common in the Palaeartic Region. Chelonus pannonicus Szépligeti, (in Budapest): Mt. Parnos, Geroplatamos, 21 May 1981, leg. I. Rozner. Described from Hungary, listed its localities in Slovakia, Bohemia, Poland and Croatia (Dalmatia). New to the fauna of Greece. Chelonus podlussanyi sp. nov.: For its description see the chapter "Description of the new species". Chelonus productus Herrich-Schaeffer, 1838 (=Ch. submuticus auct. nec Wesmael, 1835) - 1 (in Budapest): Crete, Antr. Iovis, Mt. Ida, 1906, leg. L. Biró. 1 (in Lund): Crete, 2 km S Kera, 8 km S Malia alt., 900 m, swept in vegetation, 27 June 1979, leg. L. Cederholm. 1 (in München): Cyprus mer., Amathus, 3 April 1984, leg. H. Weiffenbach. 1 (in Budapest): Prov. Drama, Mt. Falakvo, 24 May 1995, leg. I. Rozner. 2 (in Lund): Rhodos, 6 km SE Laerma, 23 May 1983, leg. R. Danielsson. 1 (in Budapest): prov. Thesaloniki, Arethousa, 23 May 1995, leg. A. Orosz. 1 (in Budapest): prov. Thessaloniki, Askos, 22 May 1995, leg. A. Orosz (in Budapest): prov. Thessaloniki, Asprovalta, 15 May 1995, leg. J. Papp. 1 (in Budapest): prov. Thessaloniki, Rendina, 21 May 1995, leg. I. Rozner. In Europe a frequent Chelonus species. Nearest to Greece reported from Bulgaria (Zaykov 1978: 231) supposedly under the name Ch. submuticus Wesmael (cf. Papp 1995: ).

11 Notes fauniques de Gembloux, n 53 (2003) 31 Chelonus scabrator (Fabritius, 1793) - 1 (in Budapest): Crete, Omalos, 18 August 1906, leg. L. Biró. Widely distributed in the Palaearctic Region. Nearest to Greece reported from Bulgaria (Zaykov 1978: 231). New to the fauna of Greece. Chelonus seticornis Thomson, (in Budapest): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Hitherto known from Sweden (Papp 1997: 16), Slovakia (Čapek & Lukaš 1989: 42) and Hungary (Papp 1996: 141). New to the fauna of Greece. Chelonus subseticornis Tobias, (in Budapest): Crete, Omalos, 8 August 1906, leg. L. Biró. Distributed in the western Palaearctic Region; nearest to Greece known from Hungary (Papp 1996: 142). New to the fauna of Greece. Microchelonus caucasicus (Abdinbekova, 1969) (=M. cretensis Tobias, 1999, syn. n.) (in Budapest): Crete, Antr. Iovis, Mt. Ida, 1906, leg. L. Biró. 1 (in Budapest): Crete, Herakleion, 5 May 1906, leg. L. Biró. 2 (in Budapest): Crete, Ins. Dia, May 1906, leg. L. Biró. 3 (in Budapest): prov. Lakonia, Mts Tagetos, m, 14 June 1979, leg. L. Gozmány. 1 (in Budapest): prov. Rethymno, Sklavokanbos, 12 May 1993, leg. A. Orosz. 1 (in Lund): Trikala, Panagia, 15 June 1982, leg. R. Danielsson (in Budapest): prov. Thessaloniki, Asprovalta, 10 May (1 + 1 ) and 15 May (1 ) 1995, leg. J. Papp. In 1999 Tobias has described the species Microchelonus cretensis, its female holotype is deposited in the Hungarian Natural History Museum, Budapest. An examination of the holotype of this species actuate me to consider this taxon as a variety of the nominate form: M. caucasicus var. cretensis (Tobias) syn. and comb. n. The flagellomeres of the female variety are conspicuously long likewise to the nominate form; the single difference between the two forms restricts to the outline (or length) of the carapace: (1) carapace of the nominate form is somewhat stouter, in dorsal view 1.75 times as long as broad, its sculpture a bit stronger (2) carapace of the variety is somewhat less stout, in dorsal view twice as long as broad, its sculpture a bit less strong. Distributed in southern half of Europe, northwards sporadic. New to the fauna of Greece. Microchelonus contractus (Nees, 1816) - 2 (in Budapest): Crete, Ins. Dia, May 1906, leg. L. Biró. 1 (in Lund): Evritania, Mt. Timfristos, 1 km S Ski Centre, 10 June 1982, leg. R. Danielsson. 1 (in Budapest): prov. Halkidiki, Mt. Holomoudas, Taxiarhis, 22 May 1995, leg. I. Rozner. 2 (in Budapest): Peloponese, Monemvasia, 15 (1 ) and 17 (1 ) April 1978, leg. J. Papp. 1 (in Lund): Trikala, Panagia, 15 June 1982, leg. R. Danielsson. A frequent Palaearctic species. Microchelonus devius (Tobias, 1964) - 2 (in Budapest): Lesboa, Mitileni, 2 August 1959, leg. L. Gozmány. The two females were compared to the types by me, they are identical with each other. Distributed in and up to now known only from the former USSR (Tobias 1986: 323) and from the former Yugoslavia (Papp 1975: 313). New to the fauna of Greece. Microchelonus gravenhorsti (Nees, 1816) - 1 (in Budapest): Corfu, leg. Paganetti. Known in several countries of Europe, nearest to Greece from Hungary (Papp 1996: 145). New to the fauna of Greece. Microchelonus microphthalmus (Wesmael, 1838) - 1 (in Budapest): prov. Thessaloniki, Askos, 22 May 1995, leg. A. Orosz. In Europe widely distributed albeit nowhere frequent. New to the fauna of Greece.

12 32 J. Papp Microchelonus subcontractus (Abdinbekova, 1971) - 1 (in Budapest): Crete, Herakleion, 26 April 1906, leg. L. Biró. 1 (in Budapest): prov. Thessaloniki, Asprovalta, 10 May 1995, leg. J. Papp. Widely distributed and fairly frequent in the western half of the former USSR, known from Hungary too. New to the fauna of Greece. Microchelonus sulcatus (Jurine, 1807) (1 in Budapest, in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. In Europe a fairly frequent species. New to the fauna of Greece. Phanerotoma (Bracotritoma) bilinea Lyle, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Achterberg (1990: 24) reported it from nine countries of Europe, nearest to Greece from Hungary. New to the fauna of Greece. Phanerotoma (Phanerotoma) dentata (Panzer, 1805) - 5 (4 in Budapest, 1 in Lund; det. H. Zettel l988): prov. Lakonia, 5 km S of Monemvasia, 16 October 1979, leg. G. Christensen. Widely distributed in the western Palaearctic Region. First reported from Greece ("Monemvasia") by Achterberg (1990: 28). Phanerotoma (Phanerotoma) fracta Kokujev, (in Budapest): Crete, Canea, August 1906, leg. L. Biró. 2 (in Budapest): Crete, Ins. Dia, May 1906, leg. L. Biró. 1 (in Budapest): Athens, Kifissia, 1 April 1978, leg. J. Papp. Widely distributed in the western Palaearctic Region. First reported from Greece ("Lakonia, 5 km S of Monemvasia") by Achterberg (1990: 32). Phanerotoma (Phanerotoma) graeca Zettel, Female holotype + male allotype (in Budapest): prov. Lakonia, 5 km S of Monemvasia, holotype: 21 September 1979 and allotype: 20 September 1979, leg. G. Christensen. 1 male paratype (in Budapest): Prov. Lakonia, Mts Taygetos, m, taken with MVlamp, July 1979, leg. L. Gozmány et G. Christensen. The species was described from Greece and up to now known only in this country. Phanerotoma (Phanerotoma) media Shestakov, (in Budapest; det. H. Zettel): prov. Lakonia, 5 km S of Monemvasia, 14 June 1980, leg. G. Christensen. Described from Uzbeghistan, sporadic in the southern half of the western Palaearctic Region. Achterberg (1990: 50) synonymized it with the name Ph. leucobasis Kriechbaumer, 1984 (=Ph. ocularis Kohl, 1906). Phanerotoma (Bracotritoma) parva Kokujev, (in Budapest): prov. Lakonia, 5 km S of Monemvasia, September 1979, leg. G. Christensen. According to Achterberg (1990: 52) distributed in four countries (Cyprus, Greece, Mongolia, European part of Russia). Phanerotoma (Bracotritoma) permixtellae Fischer, (in Lund): Rhodos, Kattavia, Ag. Pavlos, 21 May l983, leg. R. Danielsson. Hitherto known only from Syria (locus typicus). New to the fauna of Greece. Phanerotoma (Phanerotoma) sardiana Zettel, female paratypes (in Budapest): prov. Lakonia, 5 km S of Monemvasia, 15 September 1979, leg. G. Christensen. Described from Sardegna (Italy) and Greece. Phanerotoma (Phanerotoma) soror Achterberg, (6 and 2 in Budapest in Lund): prov Lakonia, 5 k S of Monemvasia, June 1979, leg. G. Christensen. Besides Greece the species has been described from Italy and France; perhaps a Mediterranean species.

13 Notes fauniques de Gembloux, n 53 (2003) 33 Phanerotoma (Bracotritoma) tritoma (Marshall, 1898) - 1 (in Budapest): prov. Lakonia, Monemvasia, 12 September 1979, leg. G. Christensen. The specimen at hand was named by H. Zettel in 1990 with a question-mark ("?tritoma"); it represents an albanic (or yellow) form of the nominate species, body entirely yellow, only first tergite laterally brownish, antenna with 23 antennomeres. Distributed in Europe, nearest to Greece reported from Italy and Cyprus (Achterberg 1990: 61). Phanerotomella bisulcata (Herrich-Schaeffer, 1838) (=Ph. nigra Szépligeti, 1900) - 1 (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1993, leg. R. Danielsson. Distributed in Germany, Austria, Bohemia, Hungary, Moldavia (Zettel 1989: 20), a rather rare species. First reported from Greece ("Monemvasia") by Achterberg (1990: 8). CARDIOCHILINAE Cardiochiles saltator (Fabricius, 1781) - 4 (1 in Budapest, 3 in München): Alt-Korinth, Peloponese, May 1963, leg.? (in Budapest): Crete, Amari, 4 June 1906, leg. L. Biró. 2 (1 in Berlin, 1 in Budapest): Linaria, Insel Skyros, May 1926, leg. Holz. 1 (in München): Loutraki, 29 May 1963, leg. M. Schwarz. 1 (in Berlin): Ost-Kreta, Kloster, Ankárathos, Ep. Padhias, 28 May 1925, leg. A. Schulz. Widely distributed and fairly frequent in the Palaearctic Region. AGATHIDINAE Agathis fulmeki Fischer, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 une 1997, leg. R. Danielsson. According to Simbolotti & Achterberg (1999: 50) known from Austria, France, Bulgaria and Morocco. New to the fauna of Greece. Agathis fuscipennis (Zetterstedt, 1838) (=A. glabricula Thomson, 1895) - 1 (in Budapest): Ost-Kreta, Ankárathos, Pedhiás Kloster, 26 May 1925, leg. A. Schulz. Widely distributed in Europe, nearest to Greece known from Bulgaria (Simbolotti & Achterberg 1999: 53). New to the fauna of Greece. Agathis montana Shestakov, (in München): Crete, Sitia, 17 May 1963, leg. M. Schwarz. 1 (in Lund): Evritania, Mt. Timfristos, 3 km S Ski Centre, ca 1400 m, 11 June 1982, leg. R. Danielsson. 1 (in Budapest): Evritania, Mt. Timfristor, 2 km N Karpinisi, 11 June 1982, leg. R. Danielsson. 1 (in Lund): Fthiotis, Iti Oros, 4 km NNE Kaloskopi, ca 1000 m, 9 June 1982, leg. R. Danielsson. 1 (in Lund): Fokis, Giona Oros, 3 km S Lefkadition, 7 June 1982, leg. R. Danielsson. 1 (in Berlin): Insel Naxos (Kyklad), Wóthri, Ost Küste, 7 September 1925, leg. A. Schulz. 1 (in Lund): Kavala, Pangeon Oros, 3 km NW Kipia, 20 June 1982, leg. R. Danielsson. In the former USSR widely distributed and frequent (Tobias 1986: 283); westwards from the former USSR listed its few localities in Hungary. New to the fauna of Greece. Simbolotti & Achterberg (1999: 88) gave its following distribution: "Andorra, Bulgaria, England, France, Kyrgyzstan, Turkey, Yugoslavia (Serbia). The listing of Hungary and Russia by Nixon (1986) is probably based on misidentifications". Agathis semiaciculata Ivanov, (in Lund): Rhodos, 6 km SE Laerma, 23 May 1983, leg. R. Danielsson. Tobias (1976: 210, 1986: 283) reported it from Ukraine, Azerbaidzhan (Mts Caucasus) and Kazakhstan; known also from Switzerland (Simbolotti & Achterberg 1999: 111). New to the fauna of Greece.

14 34 J. Papp Agathis tibialis Nees, (in Lund): Rhodos, 5 km W Maritsa, 15 May 1983, leg. R. Danielsson. Reported its localities from The Netherland, Germany and Hungary (Simbolotti & Achterberg 1999: 120). Agathis umbellatarum Nees, (in Berlin): Insel Kéa (Tsiá Kiklade), Berggegend Paläomilos, 27 September 1925, leg. A. Schulz. 1 (in Berlin): Kreta, Canea, 6-10 June 1942, leg. K. Zimmermann. 1 (in Berlin): Ost-Kreta, Kloster Ankárathos, Ep. Pedhiás, 28 May 1925, leg. A. Schulz. 2 (1 in Berlin, 1 in Budapest): Ost-Kreta, Knossos südlich von Iraklion, 29 June 1925, leg. A. Schulz. 1 (in Berlin): Ost-Kreta, Patsides, Ep. Témenos, 29 May 1925, leg. A. Schulz.4 (3 in Berlin, 1 in Budapest): Rodi, Egeo, May 1939, leg. R. Meyer. 1 (in Lund): Trikala, Orthovonnion, 15 June 1982, leg. R. Danielsson. 1 (in Budapest): West- Kreta, Kloster Goniá an Halbinsel Spátha, Eperch. Kisamos, 8 October 1925, leg. A. Schulz. Distributed in the western Palaearctic Region. Nixon (1986: 197) reported its five localities in Greece. In Greece a frequent braconid species. Agathis varipes Thomson, (in Berlin): Ost-Kreta, Iraklion (Cándia), 3 May 1925, leg. A. Schulz. 1 (in Budapest): Ost-Kreta, Knossos, südlich von Iraklion, 29 May 1929, leg. A. Schulz. Distributed in the western Palaearctic Region. New to the fauna of Greece. Bassus cingulipes (Nees, 1814) - 1 (in Lund): Rhodos, Kattavia, Ag. Pavlos, 21 May 1983, leg. R. Danielsson. Distributed in the Palaearctic Region, in Europe frequent. New to the fauna of Greece. Bassus tumidulus (Nees, 1814) - 2 (1 in Budapest, 1 in Salzburg: Coll. M. Schwarz): Kreta, Modi, W Chania, 10 October 1993, leg. M. Schwarz. Widely distributed in the western Palaearctic Region (Simbolotti & Achterberg 1992: 42). New to the fauna of Greece. MIRACINAE Mirax rufilabris Haliday, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Distributed in the western Palaearctic Region, a less frequent to sporadic species. New to the fauna of Greece. NEONEURINAE Elasmosoma berolinense (Ruthe, 1858) - 1 (in Lund): Evritania, Mt. Timfristos, 4 km Ekarpinisi, 11 June 1982, leg. R. Danielsson. Distributed in The Netherland, Germany, Hungary, Bulgaria, Russia, Mongolia. New to the fauna of Greece. Elasmosoma platamonense Huddleston, (holotype and 2 paratypes) (1 paratype in Budapest, holotype and 1 paratype in London): Platamon, August 1964, leg. R. D. Harkness. Distribution: Egypt, Greece. ORGILINAE Orgilus pimpinellae Niezabitowski, (in Budapest): Crete, Herakelion, 5 June 1906, leg. L. Biró. 1 (in Lund): Kavala, Panageon Oros, 6 km NW Kipia, 19 June 1982, leg. R. Danielsson. 1 (in Lund): Trikala, Orthovounion, 15 June 1982, leg. R. Danielsson. Widely distributed in the Palaeactic Region (Taeger 1989: 157). Orgilus ponticus Tobias, (in Lund): Rhodos, Kattavia, Ag. Pavlos, 21 May 1983, leg. R. Danielsson. Described by Tobias (1986: 272) from the European part of Russia (Krasnodarsk region); Taeger (1989: ) reported it from several

15 Notes fauniques de Gembloux, n 53 (2003) 35 European countries (Albania, Hungary, Italy, former Yugoslavia) and Asiatic Turkey. New to the fauna of Greece. Orgilus tobiasi Taeger, (in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Widely distributed in the western Palaearctic Region (Taeger 1989: 200). New to the fauna of Greece. OPIINAE Opius (Adontopius) danielssoni sp. nov. - For its description see the chapter "Description of the new species". Opius (Opiothorax) graecus Papp, 1982, species name revalidated (= Opius pappianus Fischer, l986 syn. n.). - 1 (in Budapest): Austria, Kärnten, Rosental, 460 m, 11 June 1958, leg. Gyllensvärd. The species has been described by me from Greece in 1982 on the basis of the female holotype specimen. After the description four years later Fischer (1986: 648) placed my name in junior synonymy as "Opius (Opiothorax) graecus Papp.(praeocc., nec Wesmael 1894)." Looking scrupulously over Wesmael s book ( ) on the Braconidae of Belgium I did not find either indication or description of a species under the name Opius graecus. The synonymization by Fischer was initiated, certainly, on the basis of Marshall s (1894: 343) phrase "...mais sans description...o. graecus, Wesm...." which is but a nomen nudum, i.e. a name suffering want of an existing species. I consider, consequently, my name Opius graecus as the name of a valid species and Fischer s replacement name Opius pappianus is its junior synonym. Opius (Allophlebus) pulchricornis Szépligeti, (in Lund) + 1 (in Budapest): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Reported from several countries of Europe; nearest to Greece from Hungary (Papp 1979: 86). Opius (Agnopius) similis Szépligeti, (in Lund) + 1 (in Budapest): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Recorded from Greece (Papp 1990: 274). Opius (Opiothorax) spretus Haliday, 1837 (=O. clarus Haliday, 1837; =O. arenosus Szépligeti, 1898) (1 + 2 in Budapest, in Lund): Epirus, Párga, 45 km NW Préveza, 9-23 June 1997, leg. R. Danielsson. Frequent in Europe; new to the fauna of Greece. ALYSIINAE Aspilota anaphoretica Fischer, (in Budapest): prov. Lakonia, Monemvasia, 15 April 1978, leg. J. Papp. Distribution: Austria, Hungary; new to the fauna of Greece. Aspilota fuscicornis (Haliday, 1838) - 1 (in Budapest): Crete, Omalos, 18 August 1906, leg. L. Biró. In Europe sporadic. New to the fauna of Greece. Aspilota minima (Thomson, 1895) - 1 (in Budapest): Crete, Herakleion, 5 May 1906, leg. L. Biró. Distribution: Sweden, Hungary; new to the fauna of Greece. Dinotrema amoenidens (Fischer, 1973) - 1 (in Budapest): Athens, Mt. Lykavitos, 11 April 1931, leg. J. Fodor. 1 (in Budapest): prov. Lakonia, Mts Taygetos, 1000 m, 11 June 1980, leg. G. Christensen. Described from Austria (Tirol), known from Hungary. New to the fauna of Greece.

16 36 J. Papp Dinotrema divisum (Stelfox et Graham, 1950) - 3 (in Budapest): Crete, Canea, February-April 1906, leg. L. Biró. 1 (in Budapest): Lamia, 200 m, 16 May 1977, leg. L. Zombori. Distribution: Ireland, Great Britain, Austria, Russia, Korea; new to the fauna of Greece. Dinotrema latifemur (Fischer, 1974) - 1 (in Budapest): Crete, Ins. Dia, 3 February 1906, leg. L. Biró. Distribution: Germany, Hungary, Korea; new to the fauna of Greece. Dinotrema paucicrenis (Fischer, 1973) - 1 (in Budapest): prov. Lakonia, Mts Taygetos, Anavriti, 1000 m, May, leg.? Distribution: Austria, Hungary, Korea; new to the fauna of Greece. Dinotrema semicompressum (Stelfox et Graham, 1949) (=Aspilota parapunctatum Fischer, 1976) - 1 (in Budapest): Corfu, leg. Paganetti. 1 (in Budapest): Lamia, 200 m, 16 May 1977, leg. L. Zombori. Distribution: Ireland, Great Britain, Austria, Hungary, Korea; new to the fauna of Greece. Dinotrema significarium (Fischer, 1973) - 1 (in Budapest): Corfu, leg. Paganetti. Distribution: Austria, Hungary, Korea; new to the fauna of Greece. Dinotrema spitzzickense (Fischer, 1976) - 1 (in Budapest): prov. Attiki, Poiania, 100 m, 3 April 1978, leg. J. Papp. Distribution: Austria; new to the fauna of Greece. Dinotrema vituperatum (Fischer, 1974) - 1 (in Budapest): Crete, Canea, March 1906, leg. L. Biró. Distribution: Austria, Hungary; new to the fauna of Greece. Descriptions of the new species Abbreviations applied in the descriptions (after Achterberg 1993a: 5): Eye - OOL = shortest distance between hind ocellus and compund eye; POL = shortest distance between hind two ocelli. Fore wing venation cu-a = transverse cubito-anal vein or nervulus; m-cu = transverse medio-cubital vein or recurrent vein; r = transverse radial vein or first section of the radial vein; M + CU1 = median vein; 1A + 2A = two sections of the submedian vein; 1-CU1 and 2-CU1 = first and second sections of the discoidal vein; 1-M = basal vein; 1-R1 and 2-R1 = first and second sections of the metacarpal vein; 2-SR = first transverse cubital vein; 3-SR and 4-SR (=SR1) = second and third sections of the radial vein; 1-SR-M = first section of the cubital vein. Subfamily BRACHISTINAE Triaspis graecus sp. nov. (Figs 1-5) Material examined (1 ). - Female holotype: Greece, Epirus, Párga, 45 km NW from Préveza, 9-23 June 1997, leg. R. Danielsson. The holotype is in good condition: mounted on a pointed card, tarsi of fore and middle legs partly less visible. Holotype is deposited in the Zoological Museum, The University, Lund. Etymology: The species name graecus refers to the country (Greece) where the type locality was established. Description of the female holotype: Body 2 mm long. Antenna somewhat shorter than body and with 19 antennomeres. First flagellomere four times and penultimate flagellomere subcubic or 1.25 times as long as broad, flagellomeres gradually and slightly thickening. Head in dorsal view (Fig. 1) less transverse, 1.6 times as broad

17 Notes fauniques de Gembloux, n 53 (2003) 37 as long, temple just longer than eye, temple constricted, occiput just excavated. Ocelli small and elliptic, OOL a bit longer than POL. Clypeus 3.2 times as wide as high, its lower margin faintly convex, laterally pointed (Fig. 2). Eye in lateral view almost twice as high as wide, temple clearly one-fourth wider than eye, latter beyond eye evenly broad (Fig. 3, see arrows). Head polished. Figs. 1-9 : Figs 1-5 : Triaspis graecus sp. nov.: 1 = head in dorsal view, 2 = clypeus, 3 = head in lateral view, 4 = distal part of right fore wing, 5 = carapace in dorsal view with indication of its sculpture. Figs 6-9. Triaspis hispanicus Papp: 6 = distal part of right fore wing, 7 = clypeus, 8 = head in lateral view, 9 = head in dorsal view. Mesosoma in lateral view 1.4 times as long as high. Notaulix evenly and moderately deep, finely crenulate. Precoxal suture missing. Mesosoma polished except propodeum. Propodeum rugulose with a less distinct areola basalis. Hind femur four times as long as broad medially. Hind tibia and tarsus equal in length. Hind basitarsus as long as tarsomeres 2-3 combined. Fore wing about as long as body. Pterostigma (Fig. 4) wide, 2.2 times as long as wide and issuing r from its middle, 1-R1 almost twice as long as 2-R1; marginal cell short, 3-4-SR ending far before tip of wing (Fig. 4, see arrows). Carapace rather globose (Fig. 5), 1.45 times as long as broad at second tergite. First tergite 1.7 times as broad behind as long medially, its pair of keels short and less distinct. Second tergite transverse, 1.9 times as broad behind as long medially. Hind margin of third tergite medially without excavation. Tergites 1-2 longitudinally striate (Fig. 5), third tergite slightly less strongly striate and medially with a polished field. Ovipositor sheath long, as long as carapace and half mesosoma combined. Antenna brown, pedicel yellow, flagellomeres 1-2 darkening yellowish brown. Head blackish brown, face and frons brown. Clypeus and mandible yellowish, palpi straw yellow. Mesosoma yellow, mesoscutum and scutellum brown. Legs yellow. Carapace brown. Wings hyaline, pterostigma and veins opaque brown.

18 38 J. Papp Male and host unknown. Distribution: Greece. The new species, Triaspis graecus, is nearest to T. hispanicus Papp and the two species are distinguished by the following features: 1 (2) Pterostigma issuing r distally from its middle; 1-R1 somewhat longer than 2-R1 (Fig. 6, see arrows). Lower margin of clypeus faintly pointed (Fig. 7). Eye in lateral view as wide as temple (Fig. 8, see arrows), temple in dorsal view slightly less constricted (Fig. 9). Antenna with 25 antennomeres. Carapace not globose, 1.7 times as long as broad medially; first tergite 1.5 times as broad behind as long medially; second tergite less transverse, 1.6 times as broad behind as long medially (Fig. 19 in Papp 1999: 287). Tergites reddish yellow. Male: 2.9 mm. Spain T. hispanicus Papp, (1) Pterostigma issuing r from its middle; 1-R1 twice as long as 2-R1 (Fig. 4, see arrows). Lower margin of clypeus faintly convex and laterally pointed (Fig. 2). Temple in lateral view clearly one-fourth wider than eye (Fig. 3, see arrows), temple in dorsal view slightly more constricted (Fig. 1). Antenna with 19 antennomeres. First tergite 1.7 times as broad behind as long medially; second tergite transverse, 1.9 times as broad behind as long medially. Tergites brown. Female: 2 mm. Greece T. graecus sp. nov. Subfamily OPIINAE Opius (Adontopius) danielssoni sp. nov. (Figs 10 16) Material examined (1 ). - Male holotype: Greece, Epirus, Párga, 45 km NW from Préveza, 9-23 June 1997, leg. R. Danielsson. The holotype is in good condition: glued by the right side of the mesosome on a pointed card, every leg well visible. Holotype is deposited in the Zoological Museum, The University, Lund. Etymology: The new species is dedicated to its collector, Dr. Roy Danielsson, Swedish entomologist and persistent explorer of the entomofauna of Greece. Description of the male holotype: Body 1.8 mm long. Antenna somewhat longer than body and with 20 antennomeres. First flagellomere 3.6 times and penultimate flagellomere 2.3 times as long as broad, flagellum evenly thick. Head in dorsal view (Fig. 10) 1.8 times as broad as long, eye twice as long as temple, temple strongly rounded, occiput weakly excavated. Ocelli elliptic, OOL twice as long as POL. Eye in lateral view 1.6 times as high as wide, temple ventrally somewhat narrowing and eye 1.4 times as wide as temple (Fig. 11, see arrows). Face twice as wide as high. Clypeus four times as wide as high (Fig. 12). Mandible gradually broadening basally (Fig. 13). Mouth open. Head polished. Mesosoma in lateral view 1.3 times as long as high. Notaulix faintly distinct and restricted to fore declivous part of mesoscutum. Mesoscutal dimple and precoxal suture absent. Mesosoma polished. Hind femur 4.2 times as long as broad medially (Fig. 14). Hind tibia one-fifth longer than hind tarsus. Hind basitarsus as long as tarsomeres 3-4 combined.

19 Notes fauniques de Gembloux, n 53 (2003) 39 Fore wing 1.3 times as long as body. Pterostigma (Fig. 15) 6.6 times as long as wide and issuing r proximally from its middle, 3 SR 1.6 times as long as 2-SR, 4-SR three times as long as 3-SR, feebly sinuate and reaching tip of wing. Subdiscal cell closed distally. Figs Opius (Adontopius) danielssoni sp. nov.: 10 = head in dorsal view, 11 = head in lateral view, 12 = clypeus, 13 = left mandible, 14 = hind femur, 15 =distal part of right fore wing, 16 = tergites 1-3. First tergite (Fig. 16) 1.2 times as long as broad behind and 1.8 times as broad as basally, pair of converging keels reaching middle of tergite, pair of spiracles somewhat beyond middle of tergite, tergite laterally and posteriorly uneven otherwise together with further tergites polished. Second tergite along its hind imaginary transverse margin just uneven (Fig. 16). Ground colour of body yellow. Scape yellow, apically together with pedicel brownish, flagellum dark brown. Mandible straw yellow, apically brown; palpi also straw yellow. Ocellar field brown. Mesoscutum very faintly reddish yellow. Hind margin of axille and metanotum brown. Hind margin of tergites 3-4 dark, tergite 5 widely antero-medially and tergite 6 almost entirely brown. Tergites 7-8 white. Legs light yellow. Wings subhyaline, pterostigma and veins greyish brownish. Female and host unknown. Distribution: Greece. The new species, Opius (Adontopius) danielssoni, is nearest to O. (A.) nanosoma Fischer and A. (O.) tenfanus Fischer, the latter two species are known to me on the basis of their description. The three species are distinguished as follows: 1 (2) First tergite longitudinally strio-rugulose, 1.6 times broader behind than basally. First flagellomere 1.25 times as long as second flagellomere. Eye in lateral view 1.8 times as high as wide and 1.2 times wider than temple (Abb. 17 in Fischer 1988/89: 90), temple evenly wide beyond eye. Clypeus five times as wide as high. Fore wing: 4-SR 2.5 times as long as 3-SR. Body black with yellow pattern. Female: 1.5 mm. Slovakia. O. (A.) nanosoma Fischer, (1) First tergite smooth and shiny, latero-posteriorly uneven, 1.8 times broader behind than basally (Fig. 16). Flagellomeres 1 and 2 equal in length. Eye in

20 40 J. Papp lateral view 1.6 times as high as wide and 1.4 times wider than temple, temple ventrally somewhat narrowing (Fig. 11). Clypeus four times as wide as high (Fig. 12). Fore wing: 4-SR three times as long as 3-SR (Fig. 15). Body yellow to light brownish yellow with a few brown pattern. Male: 1.8 mm. Greece O. (A.) danielssoni sp. nov. 1 (2) Eye in dorsal view less long, 1.2 times as long as temple. Antenna with antennomeres. Hind femur six times as long as broad. Fore wing: 4-SR 2.4 times as long as 3-SR. First tergite 1.5 times broader behind than basally, 1.3 times as long as broad behind. Body black with yellow pattern. Male: 2 mm. Spain: Canary Islands O. (A.) tenfanus Fischer, (1) Eye in dorsal view long, twice as long as temple (Fig. 10). Antenna with 20 antennomeres. Hind femur 4.2 times as long as broad (Fig. 14). Fore wing: 4- SR three times as long as 3-SR (Fig. 15). First tergite 1.8 times broader behind than basally, 1.2 times as long as broad behind (Fig. 16). Body yellow to light brownish yellow with a few brown pattern. Male: 1.8 mm. Greece O. (A.) danielssoni sp. n. Taxonomic remark. - The subgenus Adontopius was described by Fischer in 1984 on the basis of his species which have been described four years earlier (1980) also by him under the name Opius (Opiothorax) addentatus. Up to now the following six Opius species are assigned to this subgenus: Opius (Adontopius) adentatus Fischer, Austria Opius (Adontopius) danielssoni sp. n. - Greece Opius (Adontopius) flavipartibus Fischer, Ukraine Opius (Adontopius) nanosoma Fischer, Slovakia Opius (Adontopius) piloralis Fischer, Slovakia, Israel Opius (Adontopius) tenfanus Fischer, Spain: Canary Islands. Subfamily CHELONINAE Chelonus gozmanyi sp. nov. (Figs 17 23) Material examined (5 ). - Female holotype and 1 female paratype: Greece, prov. Lakonia, Monemvasia, 2 June 1979, leg. L. Gozmány. 3 female paratypes: Switzerland, Kanton Graubünden, Engadin, Val Roseg, 9 August 1964 (1 ) and 20 August 1964 (1 ), 1 : Pontresina 13 August 1964, leg. J. Erdõs. Holotype and four paratypes are deposited in the Hungarian Natural History Museum (Department of Zoology), Budapest, Hym. Typ. Nos (holotype) and (paratypes). The female holotype is in good condition: (1) pinned by middle of the mesosoma, (2) left flagellum damaged. The four female paratypes are also in good condition: (1) every specimen pinned by middle of the mesosoma, (2) female of No missing right fore leg except coxa + trochanter, (3) left fore wing of the female of No somewhat damaged distally beyond pterostigma, (4) distal third of left fore wing of the female No somewhat shrivelled.

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