All diving beetle specific and subspecific names explained

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1 All diving beetle specific and subspecific names explained by Anders N. Nilsson NEF > SUPPLEMENT 1, 2010 < NEF

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3 Skörvnöpparn, Umeå Supplement : 1-42 All diving beetle specific and subspecific names explained ANDERS. N. NILSSON Introduction The scientific name a of species is a binomen formed from a combination of the genus name and the specific name. To the name of a subspecies is added also a subspecific name. Specific and subspecific names are coordinated in zoology, i.e. all names within the so called species-group are available for usage at both levels (ICZN 1999: Article 46). The formation and usage of speciesgroup names is regulated by the Code (ICZN 1999), and the meaning of names is from this perspective of some interest. As species-group names have to agree in gender with the generic names which they are combined with, it has to be known if, and in what way, a certain name has to be inflected or not. One should here note, however, that my interpretations of inflections, as given in the list below, can always be challenged. Many names may be regarded as noun or adjectives, and according to the Code (Article ) the correct interpretation depends on the evidence given in the original description itself. As most older, and quite a few of the modern names were given without clear etymological statements, their intented meanings in many cases remain ambiguous. The meaning of names is of course also of some interest in itself, providing some insight into the minds and activities of those who have described the species and chosen their names. As an offspring from my work with the compilation of a world catalogue of the diving beetles in the family Dytiscidae (Nilsson 2001), I have earlier presented some more statistical aspects on the meaning and structure of specific and subspecific diving beetle names (Nilsson 2008). One of the more interesting results of that study being the strong tendency for patronyms and names based on geographical units being preferred by modern authors, instead of the more descriptive names in favour of the classical authors. A more linguistic interest for names is apt to create an awareness of and dislike for misspellings and other ill-formed names. One could argue that correctly formed and spelled names would be easier to remember and interpret for those who handle them. But the present trend in the interpretation of the Code seems to favour the usage of original spellings in most situations, nonwithstanding the missing correspondence between their formulation and intended meanings. Since October 2009 I do not any longer view the correction of a suffix of a species-group name formed from a personal name as being mandatory (cf. Article of the Code). Consequently, the original spellings of such names are now used as valid in my world catalogue, whatever etymological information was given in the original description, and e.g. Hydroporus rubyae is now considered an unjustified emendation of Hydroporus rubyi Larson, 1975, and both names are given in the list below. The present interpretation of the Code is based on the inquiry to the ICZN office made by Dr. H. Fery in This also means that all my corrections of patronyms published in Nilsson (2007) now have to be vieved as unjustified emendations, and the corresponding original spellings as the valid names of the affected species. I really feel sorry for having produced these not needed new names. There is a long tradition for providing explanations of species names in coleopterology, and I have found Schenkling s (1922) work especially inspiring. Much time has passed, however, since Schenkling compiled his list, representing only the German fauna as known by then, and with the present list I would like to expand and update the explanation of names given to species and subspecies associated with the family Dytiscidae up to the present day. As my list most probably includes several misinterpreations on the meanings of names in

4 2 Anders N. Nilsson combination with some suboptimal translations, the reader is encouraged to communicate all suggestions of improvements directly to the author. One should also remember that meanings are to be found at different levels: the literal meaning, the intended meaning, the hidden meaning, and so on. I have not separated these levels in a strict way, although I am aware of the problem. How to read the name list Each entry is given as: name in bold, genus endings (if present), explanation (meaning, person, geographical unit etc.), name kind (abbreviated), language (optional), and word class (abbreviated). Gender endings of adjectives are given in the following order: (1) masculine, (2) feminine, and (3) neuter. When more than one ending have been used, only one of the forms is listed as an entry, giving priority to masculinum. Name kind categorized as: (comb.) combination, (descr.) descriptive, (epoch) epoch or time period, (geogr.) geographical name, (hab.) habitat, (myth.) mythological, (other) other, (pers.) personal name (* indicates female), (people) people or tribe, and (?) unknown. Word class abbreviated as: (adj. comp.) comparative adjective, (adj. ) inflected adjective, (adj. name) name as adjective, (adj. un) uninflected adjective, (adj. ) as adjective (e.g. perfect and present participle), (noun app.) noun in apposition, (noun gen.) noun in genitive form, and (prep.) preposition. Identical names with different meanings are separated by their different original genus names. All mandatory changes of spelling of names have been applied, i.e. not all names are given in their original form. All known names are included, valid or invalid, available or not. Names given to fossil species are also included. When a name is interpreted as misspelled, I have added the supposed correct spelling within parenthesis after the explanation. Alphabetical name list abbreviatus, us, a, um: shortened (descr., Latin), adj. abdita, us, a, um: concealed (descr., Latin), adj. abdominalis, is, is, e: pertaining to abdomen (descr., Latin), adj. aberrans: deviating (descr., Latin), adj. aberratus, us, a, um: deviated (descr., Latin), adj. abessinicus, us, a, um: Abessinia (Ethiopia), (geogr.), adj. name abjectus, us, a, um: cast aside (descr., Latin), adj. ablutus, us, a, um: washed away (descr., Latin), adj. abnormicollis, is, is, e: with abnormal neck (descr., Latin), adj. abnormipenis: abnormal penis (descr., Latin), noun app. abonnenci: Abonnenc (pers.), noun gen. abotti: Abott (pers.), noun gen. abraeoides: Abraeus-like (descr.), adj. un absconditus, us, a, um: hidden (hab., Latin), adj. absonus, us, a, um: incompatible (descr., Latin), adj. abusus, us, a, um: misused (descr., Latin), adj. abyssinicus, us, a, um: Abyssinia (Ethiopia), (geogr.), adj. name acadianus, us, a, um: Acadia (Nova Scotia), (geogr.), adj. name acamas: son of Theseus and Phaedra (myth., Greek), noun app. acaroides: Acari-like (descr.), adj. un acharistus, us, a, um: ungracious, unpleasant (descr., Greek), adj. aciculatus, us, a, um: scratched with needle (descr., Latin), adj. acollensis, is, is, e: Acolla (geogr.), adj. name acuductus: acute formation (descr., Latin), noun app. aculeata, us, a, um: with sting (descr., Latin), adj. acuminatellus, us, a, um: with small point (descr., Latin), adj. acuminatus, us, a, um: with point that stinged (descr., Latin), adj. acutangulus, us, a, um: sharp-angled (descr., Latin), adj. acutipenis: sharp penis (descr., Latin), noun app. acutus, us, a, um: pointed (descr., Latin), adj. adachii: T. Adachi (pers.), noun gen. adamsii: Ernest Adams (pers.), noun gen. addendus, us, a, um: that ought to be added (descr., Latin), adj. adelardi: Adelard Lucien Rochette (pers.), noun gen. adelphus: brother (descr., Greek), noun app. adipatus, us, a, um: fattened (descr., Latin), adj. adjutor: supporter (descr., Latin), noun app. admotus, us, a, um: situated near (descr., Latin), adj. adpressus, us, a, um: pressed (descr., Latin), adj. adspersus, us, a, um: sprinkled (descr., Latin), adj. adumbratus, us, a, um: vague, sketchy (descr., Latin), adj. adustus, us, a, um: burnt (descr., Latin), adj. advena: a recently arrived (descr., Latin), noun app. aegyptiacus, us, a, um: Egypt (geogr., Latin), adj. name aemulus, us, a, um: competing (descr., Latin), adj. aeneolus, us, a, um: with weak tinge of copper or bronze (descr., Latin), adj. aenescens: copper-tinged (descr., Latin), adj. aeneus, us, a, um: with tinge of copper or bronze (descr., Latin), adj. aenigmaticus, us, a, um: enigmatic (descr., Latin), adj. aequabilis, is, is, e: even (descr., Latin), adj. aequalis, is, is, e: similar (descr., Latin), adj. aequatorialis, is, is, e: from equator (hab., Latin), adj.

5 All diving beetle specific and subspecific names explained 3 aequatorius, us, a, um: from equator (hab., Latin), adj. aequimarginatus, us, a, um: with identical margins (descr., Latin), adj. aequinoctialis, is, is, e: occuring when day and night are of same duration (hab., Latin), adj. aequus, us, a, um: even, flat (descr., Latin), adj. aeratus, us, a, um: with tinge of copper or bronze (descr., Latin), adj. aereus, us, a, um: with tinge of copper or bronze (descr., Latin), adj. aeruginosus, us, a, um: covered by verdigris (descr., Latin), adj. aethiopicus, us, a, um: Ethiopia (geogr.), adj. name affinis, is, is, e: closely related (descr., Latin), adj. afflatus: breath (other, Latin), noun app. afghanicus, us, a, um: Afghanistan (geogr.), adj. name africanus, us, a, um: Africa (geogr.), adj. name agaboides: Agabus-like (descr.), adj. un agaroensis, is, is, e: Agaro (geogr.), adj. name agassizi: Louis Rodolphe Agassiz (pers.), noun gen. agathodaemon: god-like demon (myth., Greek), noun app. agilis, is, is, e: agile, busy (descr., Latin), adj. agnitus, us, a, um: recognized (descr., Latin), adj. agnus: lamb (= Schäflein in German; dedicated to H. Schaeflein; also anagram of R. Angus), (other, Latin), noun app. agnusi: Colonel A. Agnus (pers.), noun gen. agrias: wild (descr., Greek), adj. un ahlwarthi: Karl Ahlwarth (pers.), noun gen. ahmeti: Ahmet Kasapoğlu (pers.), noun gen. aipo: Aipo people of New Guinea (people), noun app. aipomek: Aipomek (geogr.), noun app. airumlus, us, a, um: Airum (geogr.), adj. name ajax: hero from Trojan war (myth., Greek), noun app. akitai: Katsumi Akita (pers.), noun gen. alaskanus, us, a, um: Alaska (geogr.), adj. name alastairi: Alastair Watts (pers.), noun gen. albarracinensis, is, is, e: Sierra de Albarracin (geogr.), adj. name albertianus, us, a, um: Albert NP (geogr.), adj. name alberticus, us, a, um: Albert NP (geogr.), adj. name albidus, us, a, um: whitish (descr., Latin), adj. albionicus, us, a, um: Albion (England), (geogr.), adj. name aldabricus, us, a, um: Aldabra (geogr.), adj. name aldessa: Aldessa (pers.), noun app. alei: Ale (pers.), noun gen. alemon: father of Myscelus (myth., Greek), noun app. alexandrae: *Alexandra Ali Cieslak (pers.), noun gen. alfredi: Alfred Ekwaghe (pers.), noun gen. [Hyphydrus] alfredi: Alfredo Borelli (pers.), noun gen. [Thermonectus] algibensis, is, is, e: Sierra de Algibe (geogr.), adj. name alhambrae: Alhambra (geogr.), noun gen. alienus, us, a, um: strange (descr., Latin), adj. alinae: Sikhote Alin (geogr.), noun gen. alleghenianus, us, a, um: Alleghenian section (geogr.), adj. name alligator: like alligator (other, Latin), noun app. alligatorensis, is, is, e: Alligator river (geogr.), adj. name alluaudi: Charles Alluaud (pers.), noun gen. alogus, is, is, e: silent (without speech), (descr., Greek), adj. alpestris, is, is, e: pertaining to Alps (hab., Latin), adj. alpicola: living in Alps (hab., Latin), noun app. alpinus, us, a, um: alpine (hab., Latin), adj. altaicus, us, a, um: Altai (geogr.), adj. name alternans: changing (descr., Latin), adj. alternatus, us, a, um: changed (descr., Latin), adj. alticola: living at high altitude (hab., Latin), noun app. alutaceus, us, a, um: like leather (descr., Latin), adj. alvarengai: Moacir Alvarenga (pers.), noun gen. alvei: of river channel (hab., Latin), noun gen. amabilis, is, is, e: lovely (descr., Latin), adj. amae: *Amy Beth M. Miller (pers.), noun gen. amaenus, us, a, um: beautiful (descr., Latin), adj. amamiensis, is, is, e: Amami-Ôshima (geogr.), adj. name amandus, us, a, um: removed (descr., Latin), adj. amaroides: Amara-like (descr.), adj. un amatolensis, is, is, e: Amatole mountains (geogr.), adj. name amazonensis, is, is, e: Amazonas (geogr.), adj. name amazonicus, us, a, um: Amazonas (geogr.), adj. name ambiguus, us, a, um: ambiguous (descr., Latin), adj. ambonensis, is, is, e: Ambon island (geogr.), adj. name ambulator: one who wanders (descr., Latin), noun app. ameliae: *Ameli (pers.), noun gen. americanus, us, a, um: America (geogr.), adj. name amguemensis, is, is, e: Amguema river (geogr.), adj. name amicorum: friends (Fery, Pederzani & Wewalka), (people, Latin), noun gen. amicus, us, a, um: friendly, lovely (descr., Latin), adj. amnicola: one that lives in river (hab., Latin), noun app. amoenus, us, a, um: beautiful (descr., Latin), adj. ampliatus, us, a, um: widened (descr., Latin), adj. ampliceps: with head wide (descr., Latin), adj. un amplicollis, is, is, e: with neck wide (descr., Latin), adj. amplicornis, is, is, e: with antennae wide (descr., Latin), adj. amplimaculatus, us, a, um: with spots wide (descr., Latin), adj. amplissimus, us, a, um: very wide (descr., Latin), adj. comp. amplus, us, a, um: spacy (descr., Latin), adj. amurensis, is, is, e: Amur (geogr.), adj. name amyae: *Amy Beth M. Miller (pers.), noun gen. amybethae: *Amy Beth M. Miller (pers.), noun gen. amydrus, us, a, um: dim, faint, obscure (descr., Greek), adj. analis, is, is, e: pertaining to abdomen (or anus), (descr., Latin), adj. anasinus, us, a, um: duck-like (descr., Latin), adj. anastomosans: connecting branching parts (descr., Greek), adj. anastomozans: connecting branching parts (descr., Greek), adj. anatolicus, us, a, um: Anatolia (geogr.), adj. name anbista:? anchoralis, is, is, e: anchored (descr., Latin), adj. ancillus, us, a, um: shielded (descr., Latin), adj. andalusiae: Andalusia (geogr.), noun gen. andamanicus, us, a, um: Andaman Islands (geogr.), adj. name andinus, us, a, um: Andes (geogr.), adj. name andobonicus, us, a, um: Andobo (geogr.), adj. name andreas: Andreas Zwick (pers.), noun app. andreinii: Alfredo Andreini (pers.), noun gen. andrewesi: Andrewes Bequest (geogr.), noun gen. [Copelatus] andrewesi: Andrewes (pers.), noun gen. [Lacconectus]

6 4 Anders N. Nilsson andrewesi: Henry Leslie Andrewes (pers.), noun gen. [Peschetius] androginus, us, a, um: female-like (descr., Greek), adj. andromorphus, us, a, um: male-like (descr., Greek), adj. angelinii: Fernando Angelini (pers.), noun gen. anggi: Anggi lake (geogr.), noun app. angolensis, is, is, e: Angola (geogr.), adj. name angularis, is, is, e: angulate (descr., Latin), adj. angulicollis, is, is, e: with neck angled (descr., Latin), adj. angulifer, er, era, erum: that carries angles (descr., Latin), adj. angulipennis, is, is, e: with wings angled (descr., Latin), adj. angulolineatus, us, a, um: with angled lines (descr., Latin), adj. angusi: Robert Angus (pers.), noun gen. angustatus, us, a, um: narrowed (descr., Latin), adj. angusticollis, is, is, e: with neck narrowed (descr., Latin), adj. angusticornis, is, is, e: with antennae narrowed (descr., Latin), adj. angustior: narrower (descr., Latin), adj. comp. angustissimus, us, a, um: very narrow (descr., Latin), adj. comp. angustulus, us, a, um: somewhat narrow (descr., Latin), adj. angustus, us, a, um: narrow (descr., Latin), adj. anisonychus, us, a, um: unequal claws (descr., Greek), adj. anitae: *Anita (pers.), noun gen. anjae: *Anja Kholin & *Anja Nilsson (pers.), noun gen. anjarum: *Anja Kholin & *Anja Nilsson (pers.), noun gen. annae: *Annette Annie Miller (pers.), noun gen. annamita: Annam (geogr.), noun gen. annectens: binding together (descr., Latin), adj. annettae: *Annett Lohse (pers.), noun gen. annobomensis, is, is, e: Anno Bom Island (Annobón), (geogr.), adj. name annulatus, us, a, um: ringed (descr., Latin), adj. annulicornis, is, is, e: with antennae ringed (descr., Latin), adj. anophthalmus, us, a, um: without eyes (descr., Greek), adj. antarcticus, us, a, um: Antarctica (geogr.), adj. name antennatus, us, a, um: with antenna (descr., Latin), adj. anthracinus, us, a, um: coal-like (descr., Latin), adj. anticatus, us, a, um: placed first (descr., Latin), adj. anticus, us, a, um: anterior (descr., Latin), adj. antidotus: antidote (descr., Latin), noun app. antiquus, us, a, um: old (other, Latin), adj. antoni: Anton Fery (grandson of H. Fery), (pers.), noun gen. antrias: cave (hab., Latin), noun app. anxius, us, a, um: anxious (descr., Latin), adj. apache: Apache people of south US (people), noun app. apalodes: slightly soft (descr., Latin), adj. un apenninus, us, a, um: Appenine mountains (geogr.), adj. name aper: unmodified (descr., Greek), adj. un apertus, us, a, um: open, exposed (descr., Latin), adj. apfelbecki: Victor Apfelbeck (pers.), noun gen. aphroditae: goddess of love (myth., Greek), noun gen. aphrodite: goddess of love (myth., Greek), noun app. aphronoscelus: silly leg (descr., Greek), noun app. apiatus, us, a, um: crisped (like a parsley leaf), (descr., Greek), adj. [Guignot 1961: 836 not apicatus] apicalis, is, is, e: relating to the tip (descr., Latin), adj. apicatus, us, a, um: pointed (descr., Latin), adj. apicicornis, is, is, e: with antennae pointed (descr., Latin), adj. apicidens: becoming pointed (descr., Latin), adj. appalachius, us, a, um: Appalachian mountains (geogr.), adj. name approximatus, us, a, um: set close (descr., Latin), adj. apure: Apure city (geogr.), noun app. apuzzoi: Stefano Apuzzo (pers.), noun gen. aquaticus, us, a, um: living in water (hab., Latin), adj. aquilonaris: the northern one (hab., Latin), noun app. aquilus, us, a, um: dark (descr., Latin), adj. arabicus, us, a, um: Arabia (geogr.), adj. name arachnoides: spider-like (descr., Greek), adj. un arauco: Arauco (geogr.), noun app. arcticus, us, a, um: northern (hab., Greek), adj. arctus, us, a, um: compressed (descr., Latin), adj. arcuatus, us, a, um: curved as a bow (descr., Latin), adj. arcufer, er, era, erum: carrying bow (descr., Latin), adj. ardeaticus, us, a, um: Ardea (geogr.), adj. name areolatus, us, a, um: reticulated (descr., Latin), adj. arevaloi: D. Celso Arévalo (pers.), noun gen. argaensis, is, is, e: Serra de Arga (geogr.), adj. name argentina: Argentina (geogr.), noun app. argentinus, us, a, um: Argentina (geogr.), adj. name aristatus, us, a, um: bristled (descr., Latin), adj. aristidis, is, is, e: Aristide Letourneux (pers.), adj. name arizonicus, us, a, um: Arizona (geogr.), adj. name armatus, us, a, um: armed (descr., Latin), adj. armeniacus, us, a, um: Armenia (geogr.), adj. name armenus, us, a, um: Armenia (geogr.), adj. name armstrongi: J. Armstrong (pers.), noun gen. arnecornelii: Arne Cornelius (pers.), noun gen. artensis, is, is, e: Art island (geogr.), adj. name artus, us, a, um: dense, compact (descr., Latin), adj. artvinensis, is, is, e: Artvin (geogr.), adj. name aruensis, is, is, e: Aru Islands (geogr.), adj. name arunachal: Arunachal Pradesh (geogr.), noun app. aruspex: personal name (myth., Latin), noun app. ascendens: ascending (descr., Latin), adj. asemus, us, a, um: signless (descr., Greek), adj. ashworthii: Ashworth (pers.), noun gen. asiaticus, us, a, um: Asia (geogr., Greek), adj. name asidytus: mud-diver (descr., Greek), noun app. askalensis, is, is, e: AÃkale (geogr.), adj. name aspera, er, era, erum: roughened (descr., Latin), adj. assalyi: René Bassin Assaly (pers.), noun gen. assamensis, is, is, e: Assam (geogr.), adj. name assimilis, is, is, e: similar (descr., Latin), adj. assinicus, us, a, um: Assinie (geogr.), adj. name astrapus, us, a, um: like lightning (descr., Greek), adj. astrictovittatus, us, a, um: with narrowed stripe (descr., Latin), adj. astrophallus: star-like penis (descr., Greek), noun app. astur: Asturia (geogr.), adj. un asymmetricus, us, a, um: asymmetrical (descr., Latin), adj. atavus: ancestor (descr., Latin), noun app. ateles: imperfect (descr., Greek), adj. un ater, er, ra, rum: black (descr., Latin), adj. aterrimus, us, a, um: very black (descr., Latin), adj. comp.

7 All diving beetle specific and subspecific names explained 5 athabascae: of where there are reeds (hab., Cree indian), noun gen. atlantis, is, is, e: Atlas mountains (geogr.), adj. name atomarius, us, a, um: undivided (descr., Latin), adj. atomus, us, a, um: very small (descr., Latin), adj. atowaso: *Atowaso Sagata (pers.), noun app. atramentarius, us, a, um: blackish (descr., Latin), adj. atratus, us, a, um: cloth in black (descr., Latin), adj. atriceps: with head black (descr., Latin), adj. un atricolor: black (descr., Latin), adj. atripennis, is, is, e: with wings black (descr., Latin), adj. atristernalis, is, is, e: with breast black (descr., Latin), adj. atropos: without return (other, Greek), noun app. atrosulcatus, us, a, um: black furrowed (descr., Latin), adj. attenuatus, us, a, um: narrowed to a point (descr., Latin), adj. atypicalis, is, is, e: not typical (descr., Latin), adj. aubei: Charles Aubé (pers.), noun gen. audeni: Kenneth F. Auden (pers.), noun gen. aulicus, us, a, um: with furrow (descr., Greek), adj. aurasius, us, a, um: golden (descr., Latin), adj. aurea, us, a, um: golden (descr., Latin), adj. auriculatus, us, a, um: with small ear (descr., Latin), adj. auritus, us, a, um: eared (descr., Latin), adj. aurofasciatus, us, a, um: banded in gold (descr., Latin), adj. auropictus, us, a, um: painted in gold (descr., Latin), adj. aurora: light (descr., Latin), noun app. austinii: Austin (pers.), noun gen. australiae: Australia (geogr.), noun gen. australianus, us, a, um: Australia (geogr.), adj. name australicus, us, a, um: Australia (geogr.), adj. name australis, is, is, e: southern (hab., Latin), adj. austriacus, us, a, um: Austria (geogr.), adj. name austrocaledonicus, us, a, um: from south New Caledonia (geogr.), adj. name austrodiscors: south of discors (hab., Latin), adj. awajewi: Awajew (pers.), noun gen. avenionensis, is, is, e: Avignon (geogr.), adj. name avunculus: uncle (descr., Latin), noun app. axillaris, is, is, e: belonging to armpit (descr., Latin), adj. ayapel: Ayapel lake (geogr.), noun app. azruensis, is, is, e: Azrou (geogr.), adj. name aztec: Aztecs (people), noun app. aztecus, us, a, um: from Aztecs (people), adj. name babai: K. Baba (pers.), noun gen. babaulti: Guy Babault (pers.), noun gen. babetteae: *Babette Wilczok (pers.), noun gen. bacchusi: Mick E. Bacchus (pers.), noun gen. bacillifer, er, era, erum: carrying small rod (descr., Latin), adj. backstromi: Bäckström (pers.), noun gen. baculiformis, is, is, e: rod-shaped (descr., Latin), adj. badeni/ -ii: Johann Andre Ferdinand Baden (pers.), noun gen. baderi: Theodor Bader (pers.), noun gen. badiellus, us, a, um: slightly chestnut coloured (descr., Latin), adj. badius, us, a, um: chestnut coloured (descr., Latin), adj. baelus: missile, arrow (descr., Greek), noun app. baeri: Gustave Adolphe Baer (pers.), noun gen. baerti: Léon Baert (pers.), noun gen. baeticus, us, a, um: Baetica (Andalusia), (geogr.), adj. name bagus, us, a, um: great, (descr., Indonesian), adj. bakeri: Charles Fuller Baker (pers.), noun gen. bakewelli/ -ii: Robert Bakewell (pers.), noun gen. balabacicus, us, a, um: Balabac island (geogr.), adj. name balcanicus, us, a, um: Balcan peninsula (geogr.), adj. name balearicus, us, a, um: Balearic Islands (geogr.), adj. name balfourbrownei: John Balfour-Browne (pers.), noun gen. baliolus, us, a, um: spotted (descr., Greek), adj. balionota, us, a, um: with back spotted (descr., Greek), adj. balkei: Michael Balke (pers.), noun gen. balli: Antoine Ball (pers.), noun gen. balneator: bath attendant (hab., Latin), noun app. baloghi: J. Balogh (pers.), noun gen. balsetensis, is, is, e: Balsetum (Le Beausset), (geogr.), adj. name balzani: Louis Balzan (pers.), noun gen. bameuli: Franck Bameul (pers.), noun gen. banajai: A. Banaja (pers.), noun gen. bangalorensis, is, is, e: Bangalore (geogr.), adj. name banksii: Joseph Banks (pers.), noun gen. baolocensis, is, is, e: Bao Loc (geogr.), adj. name baoulicus, us, a, um: Baouli tribe (people), adj. name bapak: father (ref. to Manfred A. Jäch), (pers., Indonesian), noun app. baptus, us, a, um: immersed in water (hab., Latin), adj. barbarae: Santa Barbara (geogr.), noun gen. [Hydroporus] barbarae: *Barbara Lee (pers.), noun gen. [Sternopriscus] barbarensis, is, is, e: Santa Barbara (geogr.), adj. name barbarus, us, a, um: Barbaria (North Africa), (geogr.), adj. name barbouri: Thomas Barbour (pers.), noun gen. barnstoni: Barnston (pers.), noun gen. barombicus, us, a, um: Barombi Mbo lake (geogr.), adj. name barong: mythological creature of Bali (myth.), noun app. bartolozzii: Luca Bartolozzi (pers.), noun gen. barwidgeeensis, is, is, e: Barwidgee station (geogr.), adj. name barysomus, us, a, um: with heavy body (descr., Greek), adj. basalis, is, is, e: basal (hab., Latin), adj. basiclarior: basally lighter (descr., Latin), adj. comp. basicollis, is, is, e: pertaining to base of neck (descr., Latin), adj. basifasciatus, us, a, um: with basal bands (descr., Latin), adj. basilewskyi: Pierre de Basilewsky (pers.), noun gen. basillaris, is, is, e: near base (descr., Latin), adj. basinotatus, us, a, um: with basal marks (descr., Latin), adj. bassignanii: Filippo Bassignani (pers.), noun gen. bassii: Bass (pers.), noun gen. batchianensis, is, is, e: Batchian (geogr.), adj. name batekensis, is, is, e: Batéké Plateaux NP (geogr.), adj. name batesi: Henry Walter Bates (pers.), noun gen. batrachinus, us, a, um: frog-like (descr., Latin), adj. baturitiensis, is, is, e: Baturiti (geogr.), adj. name baudii: Flaminio Baudi di Selve (pers.), noun gen. baxii: Baxi (pers.), noun gen. baylyi: Ian Bayly (pers.), noun gen. bechynei: Jan Bechyné (pers.), noun gen. bedeli: Ernest Marie Louis Bedel (pers.), noun gen. bedoanus, us, a, um: Bedo (geogr.), adj. name

8 6 Anders N. Nilsson beduinus, us, a, um: Beduin brown (descr.), adj. beeri: Ernst Beer (pers.), noun gen. befasicus, us, a, um: Befasy (geogr.), adj. name behningi: A.L. Behning (pers.), noun gen. belfragei: Gustav Wilhelm Belfrage (pers.), noun gen. belli: Thomas Reid Davys Bell (pers.), noun gen. bellicosus, us, a, um: hostile (descr., Latin), adj. bellissimus, us, a, um: very beautiful (descr., Latin), adj. comp. bellitae: *Bellita (pers.), noun gen. bellus, us, a, um: beautiful (descr., Latin), adj. bengalensis, is, is, e: Bengal (geogr.), adj. name benoiti: Benoit (pers.), noun gen. benzeli: Benzel (pers.), noun gen. berdoa: Berdo (geogr.), adj. un bergeri: Berger (pers.), noun gen. bergi: Lev Semenovitch Berg (pers.), noun gen. bergmani: Sten Bergman (pers.), noun gen. berneri: Lewis Berner (pers.), noun gen. beroni: P. Beron (pers.), noun gen. bertae: *Mrs. Bertha Bert F. Carr (pers.), noun gen. bertolinii: Stefano di Bertolini (pers.), noun gen. bertrandi: Henri Bertrand (pers.), noun gen. besucheti: Claude Besuchet (pers.), noun gen. betsimisarakus, us, a, um: from land of Betsimisarakas (people), adj. name bhutanensis, is, is, e: Bhutan (geogr.), adj. name biai: Paul Bia (pers.), noun gen. bialveus, us, a, um: with 2 pits (descr., Latin), adj. bibulus, us, a, um: addicted to drinking (descr., Latin), adj. bicarinatus, us, a, um: with 2 keels (descr., Latin), adj. bicinctus, us, a, um: with 2 girdles (descr., Latin), adj. bicolor: of two colours (descr., Latin), adj. biconnatus, us, a, um: similar in two ways (descr., Latin), adj. bicornis, is, is, e: with 2 antennae (descr., Latin), adj. bicostatus, us, a, um: with 2 ribs (descr., Latin), adj. bicristata, us, a, um: with 2 crests (descr., Latin), adj. bicruciatus, us, a, um: with 2 crosses (descr., Latin), adj. bidentatus, us, a, um: with 2 teeth (descr., Latin), adj. bidessoides: Bidessus-like (descr.), adj. un bieneri: Alfred Biener (pers.), noun gen. bifarius, us, a, um: double (descr., Latin), adj. bifasciatus, us, a, um: with 2 bands (descr., Latin), adj. bifenestratus, us, a, um: with 2 windows (spots), (descr., Latin), adj. bifidus, us, a, um: split into 2 parts (descr., Latin), adj. biformis, is, is, e: with 2 forms (descr., Latin), adj. bifoveolatus, us, a, um: with 2 small pits (descr., Latin), adj. bigamus, us, a, um: of 2 fused parts (descr., Latin), adj. bigbellensis, is, is, e: Big Bell mine (geogr.), adj. name bigoti: Bigot (pers.), Latin), noun gen. biguttatus, us, a, um: with 2 spots (descr., Latin), adj. biguttulus, us, a, um: with 2 small spots (descr., Latin), adj. bihamatus, us, a, um: with 2 spines (descr., Latin), adj. biharensis, is, is, e: Bihar (geogr.), adj. name bilardoi: Armando Bilardo (pers.), noun gen. bilineatus, us, a, um: with 2 lines (descr., Latin), adj. bilita: *aunt Billy (pers.), noun app. bilix: having 2 threads (descr., Latin), adj. un bilobus, us, a, um: with 2 lobes (descr., Latin), adj. biltoni: David T. Bilton (pers.), noun gen. bilunatus, us, a, um: with 2 half moons (descr., Latin), adj. bimaculatus, us, a, um: with 2 spots (descr., Latin), adj. bimarginatus, us, a, um: with 2 borders (descr., Latin), adj. binaghii: Giovanni Binaghi (pers.), noun gen. binotatus, us, a, um: with 2 marks (descr., Latin), adj. biocellatus, us, a, um: with 2 small eyes (descr., Latin), adj. biolleyi: Biolley (pers.), noun gen. bipunctatus, us, a, um: with 2 punctures (descr., Latin), adj. bipustulatus, us, a, um: with 2 small blisters (descr., Latin), adj. biremis: ship with double oars (descr., Latin), noun app. birmanicus, us, a, um: Birma (Burma), (geogr.), adj. name biroi: Lajos (Ludwig) Biró (pers.), noun gen. bironicus, us, a, um: Lajos (Ludwig) Biró (pers.), adj. name bisbiguttatus, us, a, um: with 2 groups of 2 spots (descr., Latin), adj. biseriatus, us, a, um: with 2 rows (descr., Latin), adj. bisignatus, us, a, um: with 2 marks (descr., Latin), adj. bistriatus, us, a, um: with 2 scratches (descr., Latin), adj. bistrigatus, us, a, um: with 2 lines (descr., Latin), adj. bistrimaculatus, us, a, um: with 2 groups of 3 spots (descr., Latin), adj. bistriolatus, us, a, um: with 2 fine lines (descr., Latin), adj. bistroemi: Olof Biström (pers.), noun gen. bisulcatus, us, a, um: with 2 furrows (descr., Latin), adj. biswasi: B. Biswas (pers.), noun gen. [Platambus] biswasi: S. Biswas (pers.), noun gen. [Lacconectus] biswasi: S.K. Biswas (pers.), noun gen. [Copelatus] bitalensis, is, is, e: Bitale (geogr.), adj. name bituberculata, us, a, um: with 2 small knobs (descr., Latin), adj. biungulatus, us, a, um: with 2 claws (descr., Latin), adj. bivirgatus, us, a, um: with 2 stripes (descr., Latin), adj. bivittatus, us, a, um: with 2 bands (descr., Latin), adj. bivulnerus, us, a, um: with 2 wounds (descr., Latin), adj. bizonatus, us, a, um: with 2 broad bands (descr., Latin), adj. bjorkmanae: *Miss Frances Bjorkman (pers.), noun gen. blakeii: Blake (pers.), noun gen. blancasi: M. Fortunato Blancas Sanchez (pers.), noun gen. blanchardi/ -ii: Charles Émile Blanchard (pers.), noun gen. blancki: A. Blanck (pers.), noun gen. blandulus, us, a, um: tender (descr., Latin), adj. blandus, us, a, um: adorable (descr., Latin), adj. blatchleyi: Willis Stanley Blatchley (pers.), noun gen. blatta: moth, midge (descr., Latin), noun app. blotei: Mr Blöte (of Leyden), (pers.), noun gen. bodemeyeri: Bodo von Bodemeyer (pers.), noun gen. bodoanus, us, a, um: Bodo von Bodemeyer (pers.), adj. name boettcheri: Georg Böttcher (pers.), noun gen. bogemani: Carl Johan Bogeman (pers.), noun gen. bogemanni: Carl Johan Bogeman (pers.), noun gen. boggianii: G. Boggiani (pers.), noun gen. bogotensis, is, is, e: Bogota (geogr.), adj. name bohemani: Carl Henrik Boheman (pers.), noun gen. bohlei: Hans Wilhelm Bohle (pers.), noun gen. bokermanni: W.C.A. Bokermann (pers.), noun gen.

9 All diving beetle specific and subspecific names explained 7 boki: Gordon Bok (pers.), noun gen. bokumanus, us, a, um: Bokuma (geogr.), adj. name bolivari: Bolivar province (geogr.), noun gen. [Neobidessus] bolivari: Simón Bolivar (pers.), noun gen. [Desmopachria] bolivianus, us, a, um: Bolivia (geogr.), adj. name boliviensis, is, is, e: Bolivia (geogr.), adj. name bomansi: Bomans (pers.), noun gen. bombycinus, us, a, um: silken (descr., Latin), adj. bonariensis, is, is, e: Buenos Aires (geogr.), adj. name bonellii: F.A. Bonelli (pers.), noun gen. bonnairii: Achille Bonnaire (pers.), noun gen. bonnelli: Daniel E. Bonnell (pers.), noun gen. bonvouloiri: Victor Henry Vicomte de Bonvouloir (pers.), noun gen. boraeorum: of the gods of the north (myth., Latin), noun gen. bordoni: Carlos Bordón (pers.), noun gen. borealis, is, is, e: northern (hab., Latin), adj. boreellus, is, is, e: small northern (hab., Latin), adj. borellii: Alfredo Borelli (pers.), noun gen. boreus, us, a, um: northern (hab., Latin), adj. boristhenicus, us, a, um: Boristhenus (Dniepr), (geogr., Latin), adj. name borkuanus, us, a, um: Borkou (geogr.), adj. name borneensis, is, is, e: Borneo (geogr.), adj. name borodini: Ivan Parfenyevich Borodin (pers.), noun gen. borrei: C.F.P.A. Preudhomme de Borre (pers.), noun gen. boryslavicus, us, a, um: Borislav (geogr.), adj. name borysthenicus, us, a, um: Borysthenus (Dniepr), (geogr.), adj. name bosnicus, us, a, um: Bosnia (geogr.), adj. name bottegoi: Vittorio Bottego (pers.), noun gen. boukali: David Boukal (pers.), noun gen. boulevardi: Boulevard street (geogr.), noun gen. bouvieri: Eugenie Louis Bouvier (pers.), noun gen. bowringii: John Charles Bowring (pers.), noun gen. brachynotus, us, a, um: with back short (descr.), adj. brachypterus, us, a, um: short-winged (descr., Latin), adj. brackeli: Van Brackel (pers.), noun gen. bramardi: Bramard (pers.), noun gen. branchiatus, us, a, um: gilled (descr., Greek), adj. brancoi: Tristão Valente e Branco (pers.), noun gen. brancuccii: Michel Brancucci (pers.), noun gen. brandti: Julius von Brandt (pers.), noun gen. brannanii: Samuel Brannan (pers.), noun gen. brasiliensis, is, is, e: Brasilia (geogr.), adj. name bredoi: Joseph Anna Erich Richard Brédo (pers.), noun gen. bremondi: P. Brémond (pers.), noun gen. breweri: Charles Brewer (pers.), noun gen. brevicollis, is, is, e: with neck short (descr., Latin), adj. brevicornis, is, is, e: with antennae short (descr., Latin), adj. breviculus, us, a, um: small and short (descr., Latin), adj. brevior, or, or, us: shorter (descr., Latin), adj. comp. brevipes: short foot (descr., Latin), noun app. brevipilis, is, is, e: with hair short (descr., Latin), adj. brevis, is, is, e: short (descr., Latin), adj. brevistriga: short line (descr., Latin), noun app. brevistrigatus, us, a, um: short-lined (descr., Latin), adj. breviusculus, us, a, um: small and short (descr., Latin), adj. brincki: Per Brinck (pers.), noun gen. brivioi: Carlo Brivio (pers.), noun gen. brodei: J.S. Brode (pers.), noun gen. bromeliarum: of Bromelias (hab.), noun app. broschii: Michael Broschi Broschkowski (pers.), noun gen. browneanus: Frank Balfour-Browne (pers.), noun gen. [Bidessonotus] browneanus: John W.A.F. Balfour-Browne (pers.), noun gen. [Hydrovatus] brownei: John W.A.F. Balfour-Browne (pers.), noun gen. browni: of brown colour (descr.), noun app. [Agabus Kamiya] browni: Edvin Brown (pers.), noun gen. [Sternopriscus] browni: E.S. Brown (pers.), noun gen. [Guignotus] browni: Williamson James Brown (pers.), noun gen. [Agabus Leech, Hydroporus] bruchi: Carlos Bruch (pers.), noun gen. brucki: Emil vom Bruck (pers.), noun gen. [Hydroporus] brucki: Brück (pers.), noun gen. [Lancetes] bruesi: Brues (pers.), noun gen. brullei: Gaspard Auguste Brullé (pers.), noun gen. brumalis, is, is, e: belonging to winter (hab., Latin), adj. brunneus, us, a, um: brown (descr., Latin), adj. brunnipennis, is, is, e: with wings brown (descr., Latin), adj. brunnipes: brown foot (descr., Latin), noun app. bryanstonii: Bryanston Street (London), (other), noun gen. bryanti: Owen Bryant (pers.), noun gen. bucheti: A. Buchet (pers.), noun gen. buettikeri: W. Büttiker (pers.), noun gen. bufo: toad (descr., Latin), noun app. bula: hallo (other, Fijian), noun app. bulbus, us, a, um: onion-like (swollen antennae), (descr., Latin), adj. bulgaricus, us, a, um: Bulgaria (geogr.), adj. name bullatus, us, a, um: with bladders (descr., Latin), adj. bungei: Alexander Andrejevitsch Bunge (pers.), noun gen. buqueti: Jean Baptiste Lucien Buquet (pers.), noun gen. buresi: Ivan Josifov Bureš (pers.), noun gen. burgeoni: Louis Burgeon (pers.), noun gen. burnsi: A.N. Burns (pers.), noun gen. burrus, us, a, um: red (descr., Latin), adj. buruensis, is, is, e: Buru island (geogr.), adj. name bussleri: Heinz Bußler (pers.), noun gen. byroensis, is, is, e: Byro station (geogr.), adj. name cacozelus, us, a, um: exaggerated (descr., Latin), adj. caecus, us, a, um: blind (descr., Latin), adj. caelatipennis, is, is, e: with wings ornamented (descr., Latin), adj. caelatus, us, a, um: ornamented (descr., Latin), adj. caesus, us, a, um: cut (transverse pronotal impression), (descr., Latin), adj. caffer: inhabitant of Caffraria (people), noun app. cagrankaya: Cagrankaya (geogr.), noun app. caiaricus, us, a, um: Cairum (Cairo), (geogr.), adj. name calabricus, us, a, um: Calabria (geogr.), adj. name calaquei: Roman Calaque (colleague), (pers.), noun gen. caledoniae: [New] Caledonia (geogr.), noun app. calidus, us, a, um: warm (descr., Latin), adj. californense, is, is, e: California (geogr.), adj. name

10 8 Anders N. Nilsson californicus, us, a, um: California (geogr.), adj. name caliginosus, us, a, um: dark (descr., Latin), adj. caligosus, us, a, um: dark (descr., Latin), adj. calileguai: Calilegua NP (geogr.), noun gen. callosus, us, a, um: callous (descr., Latin), adj. calvus, us, a, um: without hair (descr., Latin), adj. cambriensis, is, is, e: Cambria (Wales), (geogr.), adj. name camerunensis, is, is, e: Camerun (geogr.), adj. name caminarius: chimney sweeper (descr., Latin), noun app. canadensis, is, is, e: Canada (geogr.), adj. name canaliculatus, us, a, um: with small furrows (descr., Latin), adj. canariensis, is, is, e: Canaria (Canary Islands), (geogr.), adj. name cancellosus, us, a, um: with grating (descr., Latin), adj. canis: dog (descr., Latin), noun app. cantabricus, us, a, um: Cantabria (geogr.), adj. name cantalicus, us, a, um: Cantal (geogr.), adj. name canthydroides: Canthydrus-like (descr.), adj. un cantralli: Irving Cantrall (pers.), noun gen. capeensis, is, is, e: Cape York (geogr.), adj. name capensis, is, is, e: Cape (geogr.), adj. name capicola: inhabitant of Cape (geogr.), noun app. capitatus, us, a, um: with head (descr., Latin), adj. capnius, us, a, um: smoky (descr., Greek), adj. caprai: Felice Capra (pers.), noun gen. capricornis, is, is, e: with antennae like goat horns (descr., Latin), adj. captiosus, us, a, um: sophisticated (descr., Latin), adj. caraboides: Carabus-like (descr.), adj. un caraibus, us, a, um: Caraibia (geogr.), adj. name carayoni: Jaques Carayon (pers.), noun gen. carbo: coal (descr., Latin), noun app. carbonarius, us, a, um: pertaining to coal (descr., Latin), adj. carbonelli: Carbonelli (pers.), noun gen. carcharias: shark (descr., Latin), noun app. cardoni: R.P. Cardon (pers.), noun gen. carinatus, us, a, um: keeled (descr., Latin), adj. carinipennis, is, is, e: with wings keeled (descr., Latin), adj. carli: Michael Carl (pers.), noun gen. carolinus: Carolina (geogr.), adj. name carrarai: Carrara (geogr.), noun gen. carri: *Bertha & John Carr (pers.), noun gen. carrorum: *Bertha & John Carr (pers.), noun gen. carstengroehni: Carsten Gröhn (pers.), noun gen. caryerus, us, a, um: like a nut (descr., Greek), adj. caspius, us, a, um: Caspia (geogr.), adj. name castaneus, us, a, um: chestnut-coloured (descr., Greek), adj. castanipennis, is, is, e: with wings chestnut-coloured (descr., Greek), adj. castilianus, us, a, um: Castilia (geogr.), adj. name casuarina, us, a, um: like casuary bird (other), adj. catascopium: scouting ship (hab., Latin), noun app. catersi: R.P.Th. de Caters (pers.), noun gen. caucasicus, us, a, um: Caucasus (geogr.), adj. name cautus, us, a, um: safe, careful (descr., Latin), adj. caviceps: with head vaulted (descr., Latin), adj. un cavicollis, is, is, e: with neck vaulted (descr., Latin), adj. cayennensis, is, is, e: Cayenne (geogr.), adj. name cazorlensis, is, is, e: Sierra de Cazorla (geogr.), adj. name cebennicus, us, a, um: Cebenna (Cévennes), (geogr.), adj. name cebuensis, is, is, e: Cebu (geogr.), adj. name celatus, us, a, um: forgotten (descr., Latin), adj. celebensis, is, is, e: Celebes (geogr.), adj. name celebesanus, us, a, um: Celebes (geogr.), adj. name celidotus, us, a, um: spotted (descr., Greek), adj. celinoides: Celina-like (descr.), adj. un celox: yacht (descr., Latin), noun app. cenchramis: fig seed (descr.), noun app. centralis, is, is, e: in middle (descr., Latin), adj. cephalotes: head-like (descr., Greek), noun app. ceramensis, is, is, e: Seram (geogr.), adj. name ceratus, us, a, um: waxy (descr., Latin), adj. cercyrae: Cercyra river nymph (myth., Greek), noun gen. ceresyi: M. de Ceresy (Lefébure de Cérisy), (pers.), noun gen. cerisyi: Lefébure de Cérisy (pers.), noun gen. cermenius: like cermeños (myth., Spanish), noun app. cervicalis: small pillow (descr., Latin), noun app. cervinus, us, a, um: deer-like (descr., Latin), adj. cervus: deer (descr., Latin), noun app. cesii: Federico Angelo Cesi (prince), (pers.), noun gen. cessatus, us, a, um: idle (descr., Latin), adj. ceylonicus, us, a, um: Ceylon (geogr.), adj. name chalconatus, us, a, um: with back earth-coloured (descr., Greek), adj. challaensis, is, is, e: Challa pastoral station (geogr.), adj. name challeti: Gilbert H. Challet (pers.), noun gen. chalybaeus, us, a, um: colour like steel (descr., Latin), adj. chalybeatus, us, a, um: shining like steel (descr., Latin), adj. championi: George Charles Champion (pers.), noun gen. [Bidessonotus, Laccophilus] championi: H.G. Champion (pers.), noun gen. [Microdytes] chapmani: Ph. Chapman (pers.), noun gen. chappuisi: P.-A. Chappuis (pers.), noun gen. charactes: engraved (descr., Greek), adj. un charaxinus, us, a, um: engraved (descr., Latin), adj. charini: V. Kharin (pers.), noun gen. charlottii: Charlotte Street (London), (other), noun gen. charon: boatman of Styx (myth., Greek), noun app. chathamicus, us, a, um: Chatham islands (geogr.), adj. name chaudoirii: Maximilien Baron de Chaudoir (pers.), noun gen. cheesmanae: *Miss L.E. Cheesman (pers.), noun gen. chei: Ernesto Che Guevara de la Serna (pers.), noun gen. chelinus, us, a, um: like turtle (descr., Greek), adj. chevrolati: Louis Alexandre Auguste Chevrolat (pers.), noun gen. chibcha: Chibcha people of Colombia (people), noun app. chibi: dwarf (descr., Japanese), noun app. chiengmaiensis, is, is, e: Chieng Mai NP (geogr.), adj. name chihuahuae: Chihuahua (geogr.), noun gen. chilensis, is, is, e: Chile (geogr.), adj. name chiliensis, is, is, e: Chili (Chile), (geogr.), adj. name chilonarius, us, a, um: pertaining to lip (descr., Greek), adj. chinensis, is, is, e: China (geogr.), adj. name chini: Chini lake (geogr.), noun app. chipi: Percival Chip Watts (pers.), noun gen.

11 All diving beetle specific and subspecific names explained 9 chipiriricus, us, a, um: Chipiriri river (geogr.), adj. name chishimanus, us, a, um: Chishima Rettô (Kuril islands), (geogr.), adj. name chloroticus, us, a, um: yellowish green (descr., Greek), adj. chopardi: Lucien Chopard (pers.), noun gen. chrisi: Chris H.S. Watts (pers.), noun gen. chrysomelinus, us, a, um: Chrysomela-like (descr.), adj. chrysostomus, us, a, um: with mouth golden (descr., Greek), adj. chujoi: Michio Chûjô (pers.), noun gen. chumphon: Chumphon province (geogr.), noun app. churchillensis, is, is, e: Churchill (geogr.), adj. name ciampori: Fedor Ciampor (pers.), noun gen. cicatricosus, us, a, um: scarred (descr., Latin), adj. cicer: chickpea (descr., Latin), noun app. cicur: tame (descr., Latin), adj. un cicurius, us, a, um: tame (descr., Latin), adj. ciliatus, us, a, um: with small hair (descr., Latin), adj. cilunculus: person with head narrow (descr., Latin), noun app. cimicioides: Cimex-like (descr.), adj. un cinctatus, us, a, um: girdled (descr., Latin), adj. cinctellus, us, a, um: with weak transverse border (descr., Latin), adj. cincticollis, is, is, e: neck with transverse band (descr.), adj. cinctipennis, is, is, e: wings with transverse band (descr.), adj. cinctus, us, a, um: with a girdle (descr., Latin), adj. cinereus, us, a, um: ash-coloured (descr., Latin), adj. cingulatus, us, a, um: with transverse band (descr., Latin), adj. cingulifer, er, era, erum: carrying transverse band (descr., Latin), adj. cinnamomeus, us, a, um: like cinnamon (descr., Latin), adj. circularis, is, is, e: rounded (descr., Latin), adj. circulatus, us, a, um: rounded (descr., Latin), adj. circumcinctus, us, a, um: circumference banded (descr., Latin), adj. circumductus: circumference (descr., Latin), noun app. circumflexus: vault (descr., Latin), noun app. circumscriptus, us, a, um: rounded (descr., Latin), adj. civicus, us, a, um: civil (descr., Latin), adj. clairvillei: Joseph Philippe de Clairville (pers.), noun gen. clarki/ -ii: Hamlet Clark (pers.), noun gen. claussi: Dr Clauss (pers.), noun gen. clavatus, us, a, um: club-like (descr., Latin), adj. clavicornis, is, is, e: with antenna clubbed (descr., Latin), adj. cleopatrae: *Cleopatra (pers.), noun gen. clessini: Clessin (pers.), noun gen. clubista:? clypealis, is, is, e: with shield (descr., Latin), adj. coarctaticollis, is, is, e: with neck compressed (descr., Latin), adj. coarcticolis, is, is, e: with neck compressed (descr., Latin), adj. coccinelloides: Coccinella-like (descr., Latin), adj. un cocheconis, is, is, e: Cocheco river (geogr.), adj. name coelamboides: Coelambus-like (descr.), adj. un coelata, us, a, um: with cavity (descr., Greek), adj. cognatus, us, a, um: similar (descr., Latin), adj. colasi: Guy Colas (pers.), noun gen. colberti: Stephen T. Colbert (pers.), noun gen. colchicus, us, a, um: Colchis (geogr.), adj. name collaris, is, is, e: pertaining to neck (descr., Latin), adj. collarti: Albert Collart (pers.), noun gen. collatus, us, a, um: condensed (descr., Latin), adj. collega: colleague (descr., Latin), noun app. collocallosus, us, a, um: neck with callus (descr., Latin), adj. coloradensis, is, is, e: Colorado (geogr.), adj. name columbianus, us, a, um: British Columbia (geogr.), adj. name colymbetoides: Colymbetes-like (descr.), adj. un colymbitoides: Colymbites-like (descr.), adj. un colymbus, us, a, um: swimming (descr., Latin), adj. comes: companion (descr., Latin), noun app. comma: comma (descr., Latin), noun app. commatifera, er, era, erum: carrying comma (descr., Latin), adj. comoensis, is, is, e: Comoé river (geogr.), adj. name compactus, us, a, um: small (descr., Latin), adj. compar: similar (descr., Latin), adj. compertus, us, a, um: investigated (descr., Latin), adj. complicatus, us, a, um: complicated (descr., Latin), adj. compotor: companion (descr., Latin), noun app. compressus, us, a, um: flat (descr., Latin), adj. compsus, us, a, um: elegant (descr., Greek), adj. comptus, us, a, um: adorned (descr., Latin), adj. compunctus, us, a, um: pricked, punctured (descr., Latin), adj. concessor: latecomer (descr., Latin), noun app. concettae: *Concetta Torregrossa (pers.), noun gen. concii: Cesare Concii (pers.), noun gen. concinnaticius, us, a, um: put together in correct way (descr., Latin), adj. concinnus, us, a, um: of pleasing proportion (descr., Latin), adj. concisus, us, a, um: short (descr., Latin), adj. concivis: a fellow-citizen (descr., Latin), noun app. concolor: of one colour (descr., Latin), adj. un concolorans: of one colour (descr., Latin), adj. un confertus, us, a, um: dense (descr., Latin), adj. confinis, is, is, e: related (descr., Latin), adj. confluens: confluence (descr., Latin), noun app. confluentus, us, a, um: confluent (descr., Latin), adj. conformis, is, is, e: shaped (descr., Latin), adj. confossus, us, a, um: with pits (descr., Latin), adj. confusus, us, a, um: confused (descr., Latin), adj. congener: relative (descr.), noun app. congeneroides: congener-like (descr.), adj. un congestus, us, a, um: crammed (descr., Latin), adj. congo: Congo (geogr.), noun app. congoanus, us, a, um: Congo (geogr.), adj. name congruus, us, a, um: corresponding (descr., Latin), adj. conicollis, is, is, e: with neck conical (descr., Latin), adj. conicus, us, a, um: conical (descr., Latin), adj. conjunctus, us, a, um: related (descr., Latin), adj. conjungens: related (descr., Latin), adj. conoideus, us, a, um: similar (descr., Latin), adj. conradsi: Hubert Aloys Conrads (pers.), noun gen. consanguineus, us, a, um: related (descr., Latin), adj. consentaneus, us, a, um: be in accordance (descr., Latin), adj.

12 10 Anders N. Nilsson consimilis, is, is, e: uniform (descr., Latin), adj. consobrinus: cousin (descr., Latin), noun app. consors: participant (descr., Latin), noun app. conspersus, us, a, um: spotted (descr., Latin), adj. conspicuus, us, a, um: conspicuous (descr., Latin), adj. consputus, us, a, um: spotted (mocked at), (descr., Latin), adj. constans: constant (descr., Latin), adj. un constantini: Robert Constantin (pers.), noun gen. consuetus, us, a, um: common (descr., Latin), adj. contiger: spear-bearer, lancer (descr., Latin), noun app. contiguus, us, a, um: adjacent (descr., Latin), adj. continentalis, is, is, e: continental (hab., Latin), adj. contiro: fellow recruit (descr., Latin), noun app. contractulus, us, a, um: contracted (descr., Latin), adj. contumax: defiant (descr., Latin), adj. un convexior: more convex (descr., Latin), adj. comp. convexiusculus, us, a, um: slightly convex (descr., Latin), adj. convexus, us, a, um: convex (descr., Latin), adj. coomani: R.P. de Cooman (pers.), noun gen. cooperae: *Joyce Omer-Cooper (pers.), noun gen. cooperi: Joseph Omer-Cooper (pers.), noun gen. [Deronectes, Herophydrus] cooperi: Steve Cooper (pers.), noun gen. [Limbodessus] copelatoides: Copelatus-like (descr.), adj. un copidotibiae: edged tibiae (descr., Latin), noun app. coquerelii: Jean Charles Coquerel (pers.), noun gen. coracinus, us, a, um: black as raven (descr., Latin), adj. corayi: Armin Coray (pers.), noun gen. cordaticollis, is, is, e: with neck heart-shaped (descr., Latin), adj. cordatus, us, a, um: heart-shaped (descr., Latin), adj. cordieri: Pierre L.A. Cordier (?), (pers.), noun gen. cordillerensis, is, is, e: Cordilleran region (hab.), adj. name cordylinoides: from Brocchinia cordylinoides (a Bromeliad), (hab.), adj. un coriacellus, us, a, um: somewhat leather-like (descr., Latin), adj. coriaceus, us, a, um: leather-like (descr., Latin), adj. cornatipenis: horned penis (descr., Latin), noun app. corniger, er, era, erum: carrying horns (descr., Latin), adj. coronatus, us, a, um: with wreath (descr., Latin), adj. corporaali: Johannes Bastiaan Corporaal (pers.), noun gen. corpulentus, us, a, um: stout (descr., Latin), adj. corsicus, us, a, um: Corsica (geogr.), adj. name corticeus, us, a, um: from bark (descr., Latin), adj. corumbensis, is, is, e: Corumba (geogr.), adj. name corvallis: Corvallis (geogr.), noun app. corvinus, us, a, um: black (descr., Latin), adj. corvus: raven (descr., Latin), noun app. corystes: helmeted, crested (descr., Greek), adj. un costalis, is, is, e: with ribs (descr., Latin), adj. costaricensis, is, is, e: Costa Rica (geogr.), adj. name costatus, us, a, um: with ribs (descr., Latin), adj. costipennis, is, is, e: with wings ribbed (descr., Latin), adj. costulatus, us, a, um: with fine ribs (descr., Latin), adj. costulifer, er, era, erum: carrying fine ribs (descr., Latin), adj. counselli: Dominic Counsell (pers.), noun gen. couragei: *Ann Courage (pers.), noun gen. couretae: *Mme. Couret (pers.), noun gen. coxalis, is, is, e: pertaining to the hip (descr., Latin), adj. crassicornis, is, is, e: with antenna robust (descr., Latin), adj. crassifrons: robust forehead (descr., Latin), noun app. crassipes: robust foot (descr., Latin), noun app. crassiusculus, us, a, um: slightly robust (descr., Latin), adj. crassulus, us, a, um: somewhat robust (descr., Latin), adj. crassus, us, a, um: robust (descr., Latin), adj. cretensis, is, is, e: Crete (geogr.), adj. name creticus, us, a, um: Crete (geogr.), adj. name cribrarius, us, a, um: with punctation dense (descr., Latin), adj. cribratellus, us, a, um: with punctation relatively dense (descr., Latin), adj. cribratus, us, a, um: densely punctate (descr., Latin), adj. cribrosus, us, a, um: with punctation dense (descr., Latin), adj. criniticoxis, is, is, e: with coxa hairy (descr., Latin), adj. crinitisternus, us, a, um: with sternum hairy (descr., Latin), adj. crispatus, us, a, um: wrinkled (descr., Latin), adj. cristatomarginatus, us, a, um: with crested margins (descr., Latin), adj. cristatus, us, a, um: crested (descr., Latin), adj. croceus, us, a, um: saffron-coloured (descr., Latin), adj. crotchi: George Robert Crotch (pers.), noun gen. crowii: Crow (pers.), noun gen. crucialis, is, is, e: with cross (descr., Latin), adj. cruciatus, us, a, um: crossed (descr., Latin), adj. crucifer, er, era, erum: carrying cross (descr., Latin), adj. cruentatus, us, a, um: made bloody (descr., Latin), adj. crux: cross (descr., Latin), noun app. cryptarchoides: Cryptarcha-like (descr.), adj. un crypticoides: Crypticus-like (descr.), adj. un crypticus, us, a, um: concealed (descr., Greek), adj. cryptolineatus, us, a, um: with lines concealed (descr., mixed), adj. cryptus, us, a, um: hidden (descr., Greek), adj. cubaensis, is, is, e: Cuba (geogr.), adj. name cubensis, is, is, e: Cuba (geogr.), adj. name cueensis, is, is, e: Old Cue bores (geogr.), adj. name cunctator: one that is late (descr., Latin), noun app. cuneatus, us, a, um: wedge-shaped (descr., Latin), adj. cunyuensis, is, is, e: Cunyu station (geogr.), adj. name cuppeni: Jan G.M. Cuppen (pers.), noun gen. cupreolus, us, a, um: with slight coppery lustre (descr.), adj. cuprescens: with coppery lustre (descr.), adj. curculinus, us, a, um: weevil-like (descr., Latin), adj. curticollis, is, is, e: with neck short (descr., Latin), adj. curticornis, is, is, e: with antenna short (descr., Latin), adj. curtistriatus, us, a, um: with shortened scratches (descr., Latin), adj. curtulus, us, a, um: short and small (descr., Latin), adj. curvifasciatus, us, a, um: with curved band (descr., Latin), adj. curvilineatus, us, a, um: with curved line (descr., Latin), adj. curvilobus, us, a, um: with median lobe curved (descr., mixed), adj. curvipes: curved foot (descr., Latin), noun app.

13 All diving beetle specific and subspecific names explained 11 curviplicatus, us, a, um: with curved fold (descr., Latin), adj. curvispinis: curved spine (descr., Latin), noun app. cuspidatus, us, a, um: pointed (descr., Latin), adj. cuspis: pointed end (descr., Latin), noun app. cyaneus, us, a, um: dark blue (descr., Latin), adj. cycloides: ring-like (descr., Latin), adj. un cyclopis, is, is, e: with eye round (descr., Latin), adj. cynthiae: *Cinzia Altieri (pers.), noun gen. cyprioticus, us, a, um: Cyprus (geogr.), adj. name cyprius, us, a, um: Cyprus (geogr.), adj. name czerskii: Alexander Ivanovitsch Cherskiy (pers.), noun gen. dabieshanensis, is, is, e: Dabie Shan (geogr.), adj. name daccordii: Mauro Daccordi (pers.), noun gen. dacunhae: Tristan da Cunha (geogr.), noun gen. daemeli: Eduard Dämel (pers.), noun gen. dagombae: Dagomba people of Ghana (people), noun gen. dahlii: Georg Dahl (pers.), noun gen. dahuricus, us, a, um: Dahuria (geogr.), adj. name daisetsuzanus, us, a, um: Daisetsu mountain (geogr.), adj. name dakarensis, is, is, e: Dakar (geogr.), adj. name dalmatinus, us, a, um: Dalmatia (geogr.), adj. name dama: buck (descr., Latin), noun app. damantiensis, is, is, e: Damanti village (geogr.), adj. name danae: *Dana (pers.), noun gen. dani: Dani people of New Guinea (people), noun app. danicus, us, a, um: Dania (Denmark), (geogr.), adj. name danielssoni: Roy Danielsson (pers.), noun gen. darfurensis, is, is, e: Darfur (geogr.), adj. name darlingtoni: Philip Jackson Jr. Darlington (pers.), noun gen. darlotensis, is, is, e: Darlot lake (geogr.), adj. name darwinii: Charles Darwin (pers.), noun gen. dasythrix: dense hair (descr., Greek), noun app. dauricus, us, a, um: Dauria (geogr.), adj. name davidis: Père Armand David (pers.), noun gen. davisii: Mr. A.H. Davis (pers.), noun gen. debarrosmachadoi: Dr. A. De Barros-Machado (pers.), noun gen. debilipes: weak foot (descr., Latin), noun app. debilis, is, is, e: incapable, weak (descr., Latin), adj. decastigma: 10 marks (descr., Latin), noun app. decellei: J. Decelle (pers.), noun gen. decemguttatus, us, a, um: with 10 spots (descr., Latin), adj. decemlineatus, us, a, um: with 10 lines (descr., Latin), adj. decemmaculatus, us, a, um: with 10 marks (descr., Latin), adj. decemnotatus, us, a, um: with 10 marks (descr., Latin), adj. decempunctatus, us, a, um: with 10 punctures (descr., Latin), adj. decemsignatus, us, a, um: with 10 marks (descr., Latin), adj. decemstriatus, us, a, um: with 10 scratches (descr., Latin), adj. decemsulcatus, us, a, um: with 10 furrows (descr., Latin), adj. deceptor: cheater (descr., Latin), noun app. deceptus, us, a, um: cheated (descr., Latin), adj. decipiens: false (descr., Latin), adj. decoratus, us, a, um: ornamented (descr., Latin), adj. decorosa, us, a, um: very fair (descr., Latin), adj. decorus, us, a, um: fair (descr., Latin), adj. definitus, us, a, um: determined (descr., Latin), adj. deflatus, us, a, um: flattened (descr., Latin), adj. deflectus, us, a, um: bent down (descr., Latin), adj. defloccata, us, a, um: bald (descr., Latin), adj. degener, er, era, erum: degenerate (descr., Latin), adj. degeneratus, us, a, um: degenerate (descr., Latin), adj. dehaanii: Dehaan (pers.), noun gen. deharvengi: Louis Deharveng (pers.), noun gen. dehraduni: Dehra Dun (geogr.), noun gen. dejeanii: P.F.M.A. Dejean (pers.), noun gen. delarouzei: Charles Delarouzée (pers.), noun gen. delectus, us, a, um: exquisite (descr., Latin), adj. deletus, us, a, um: denied (descr., Latin), adj. delfini: Federico Delfin (pers.), noun gen. delibatus, us, a, um: diminished (descr., Latin), adj. delicatulus, us, a, um: fine and small (descr., Latin), adj. delictus, us, a, um: faulted (descr., Latin), adj. delkeskampi: Kurt Delkeskamp (pers.), noun gen. deltoides: like Greek letter delta (triangular), (descr., Latin), adj. un demarcoi: Paulo de Marco Jr. (pers.), noun gen. dembickyi: Lubos Dembický (pers.), noun gen. demoflysi: R. Démoflys (pers.), noun gen. demoulini: Demoulin (pers.), noun gen. denigratus, us, a, um: totally blackened (descr., Latin), adj. denikei: W.N. Denike (pers.), noun gen. densatus, us, a, um: condensed (descr., Latin), adj. densepunctatus, us, a, um: densely punctate (descr., Latin), adj. densus, us, a, um: dense (descr., Latin), adj. dentatus, us, a, um: toothed (descr., Latin), adj. dentellus, us, a, um: with small tooth (descr., Latin), adj. denticulatus, us, a, um: toothed (descr., Latin), adj. dentiger, er, era, erum: carrying tooth (descr., Latin), adj. depauperatus, us, a, um: impoverished (descr., Latin), adj. depictus, us, a, um: painted (descr., Latin), adj. deplanatus, us, a, um: flattened (descr., Latin), adj. depressicollis, is, is, e: with neck compressed (descr., Latin), adj. depressus, us, a, um: compressed (descr., Latin), adj. derasus, us, a, um: shaved (descr., Latin), adj. derelictus, us, a, um: fraudulent (descr., Latin), adj. dermestoides: Dermestes-like (descr.), adj. un descarpentriesi: Descarpentries (pers.), noun gen. deserticola: one that lives in deserts (hab.), noun app. desertorum: of deserts (hab.), noun gen. desii: Kai Onkel Desi Desinger (pers.), noun gen. desintegratus, us, a, um: fragmented (descr.), adj. desjardinsii: Julien Fraçois Desjardins (pers.), noun gen. despectus, us, a, um: unnoticed (descr., Latin), adj. detectus, us, a, um: detected (descr.), adj. detritus: distintegrated material (descr., Latin), noun app. dettneri: Konrad Dettner (pers.), noun gen. deumaculatus, us, a, um: with 2 spots (descr.), adj. devexus, us, a, um: inclined (descr.), adj. deviata, us, a, um: deviating (descr.), adj. devillei: J. Saint Claire Deville (pers.), noun gen. devroyei: Devroy (pers.), noun gen. dhofarensis, is, is, e: Dhofar (geogr.), adj. name

14 12 Anders N. Nilsson diabolicus, us, a, um: devil-like (descr., Greek), adj. diastatus, us, a, um: segregated (descr., Greek), adj. dichrous, us, a, um: of 2 colours (descr., Greek), adj. didymoides: didymus-like (descr.), adj. un didymus, us, a, um: split in 2 halves (descr., Greek), adj. dieterlei: *Elisabeth, Klaus & Stephan Dieterle (pers.), noun gen. dieterleorum: *Elisabeth, Klaus & Stephan Dieterle (pers.), noun gen. differens: difference (descr., Latin), noun app. difficilis, is, is, e: difficult (descr., Latin), adj. diffinis, is, is, e: defined (descr., Latin), adj. diffisus, us, a, um: extended (= diffusus), (descr., Latin), adj. difformis, is, is, e: of different form (descr., Latin), adj. digitalis, is, is, e: finger-like (descr., Latin), adj. dikinohaseus: Naoto Kino, Kyoichi Kinomura & Yoh Hasegawa (2 kino + hase), (pers.), noun app. dilatatus, us, a, um: dilated (descr., Latin), adj. dilatipennis, is, is, e: with wings dilated (descr., Latin), adj. dilutus, us, a, um: diluted (descr., Latin), adj. dimidiatus, us, a, um: medially split (descr., Latin), adj. dimorphus, us, a, um: of 2 forms (descr., Latin), adj. dineutoides: Dineutus-like (descr.), adj. un dingbatensis, is, is, e: Dingbat well (geogr.), adj. name dintelmanni: Horst Dintelmann (pers.), noun gen. diruptus, us, a, um: teared apart (descr., Latin), adj. discedens: separating (descr., Latin), adj. discicollis, is, is, e: with neck disc-like (descr., Latin), adj. discifer, er, era, erum: carrying disc (descr., Latin), adj. discindens: tearing to pieces (descr., Latin), adj. discoidalis, is, is, e: disc-like (descr.), adj. discoideus, us, a, um: disc-like (descr.), adj. discolor: of different colours (descr., Latin), adj. un discors: disunited (descr., Latin), adj. un discrepatus, us, a, um: disagreeing (descr., Latin), adj. discretus, us, a, um: separated (descr., Latin), adj. disintegratus, us, a, um: fragmented (descr., Latin), adj. disjectus, us, a, um: scattered (descr., Latin), adj. disjunctus, us, a, um: separated (descr., Latin), adj. dismalus, us, a, um: Dismal swamp (geogr.), adj. name dismorphus, us, a, um: with 2 forms (descr., Greek), adj. dispar: unlike (descr., Latin), adj. un dispersus, us, a, um: spread out (descr., Latin), adj. dispositus, us, a, um: distributed (descr., Latin), adj. dissentiens: becoming secluded (descr., Latin), adj. dissimilis, is, is, e: different (descr., Latin), adj. distantus, us, a, um: remote (descr., Latin), adj. distigma: 2 marks (descr., Latin), noun app. distinctus, us, a, um: distinctive (descr., Latin), adj. distinguendus, us, a, um: that ought to be separated (descr., Latin), adj. ditylus, us, a, um: with 2 humps (descr., Greek), adj. diversicornis, is, is, e: with antennae different (descr., Latin), adj. diversipes: different foot (descr., Latin), noun app. divisus, us, a, um: parted (descr., Latin), adj. divoi: Divo (geogr.), noun gen. dixianus, us, a, um: Dixie (S USA), (geogr.), adj. name djurdjurensis, is, is, e: Djurdjura (geogr.), adj. name dobrogeanus, us, a, um: Dobrudscha (geogr.), adj. name doderoi: Agostino Dodero (pers.), noun gen. dodgei: H. Rodney Dodge (pers.), noun gen. dohrni: Carl August Dohrn (pers.), noun gen. dolabratus, us, a, um: axe-like (descr., Latin), adj. dolerosus, us, a, um: deceitful (descr., Greek), adj. dolichodactylus, us, a, um: with toe long (claws), (descr., Greek), adj. dollmani: Hereward Chune Dollman (pers.), noun gen. dolomitanus, us, a, um: Dolomiten (geogr.), adj. name dolosus, us, a, um: deceptive (descr., Latin), adj. dominguensis, is, is, e: Santo Domingo (geogr.), adj. name dongba: culture of Nathi people of Yunnan (other), noun app. doriae: Giacomo Doria (pers.), noun gen. dorsalis, is, is, e: relating to back (descr., Latin), adj. dorsiger, er, era, erum: carrying on one s back (descr., Latin), adj. dorsoplagiatus, us, a, um: dorsally warped (descr., Greek), adj. dorsovittatus, us, a, um: dorsally lined (descr.), adj. doubledayii: Doubleday (pers.), noun gen. draco: dragon (shape of genitalia), (descr., Latin), noun app. dregei: Carl Friedrich Drege (pers.), noun gen. dreheri: Dreher (pers.), noun gen. dressleri: Herrn Steuerverwalter Dressler (pers.), noun gen. drewseni: Drewsen (pers.), noun gen. drymetes: one that dwells in forest (hab., Greek), noun app. dubiosus, us, a, um: very doubtful (descr., Latin), adj. dubius, us, a, um: doubtful (descr., Latin), adj. duboisi: Albert Dubois (pers.), noun gen. ducalis, is, is, e: pertaining to duke (descr., Latin), adj. dudgeoni: David Dudgeon (pers.), noun gen. dufaui: Léo Dufau (pers.), noun gen. duftschmidi/ -ii: Caspar Erasmus Duftschmid (pers.), noun gen. duftschmidti: Caspar Erasmus Duftschmid (pers.), noun gen. duodecimlineatus, us, a, um: with 12 lines (descr., Latin), adj. duodecimmaculatus, us, a, um: with 12 spots (descr., Latin), adj. duodecimpustulatus, us, a, um: with 12 small blisters (descr., Latin), adj. duodecimstriatus, us, a, um: with 12 scratches (descr., Latin), adj. duplex: double (descr., Latin), adj. un duplicatus, us, a, um: doubled (descr., Latin), adj. duponti: Dupont (pers.), noun gen. duvergeri: Joseph-Alexandre Duverger (pers.), noun gen. dyscheres: bad heir (descr., Latin), noun app. dyseres: bad heir (descr., Latin), noun app. dytiscoides: Dytiscus-like (descr.), adj. un dzieduszyckii: Dzieduszycki (pers.), noun gen. eberhardi: Stefan Eberhard (pers.), noun gen. eboris: of ebony (descr., Latin), noun gen. ecuadorius, us, a, um: Ecuador (geogr.), adj. name edax: voracious (descr., Latin), adj. un edenianus, us, a, um: Castle Eden (geogr.), adj. name edithae: *Edith A. Miller (pers.), noun gen. edwardsi: Oliver T. Edwards (pers.), noun gen. effeminatus, us, a, um: weakened (descr., Latin), adj. efoutensis, is, is, e: Efout village (geogr.), adj. name

15 All diving beetle specific and subspecific names explained 13 eikei: Eike Hendrich (father of L. Hendrich), (pers.), noun gen. ejactus, us, a, um: ejected (descr., Latin), adj. ejuncidus, us, a, um: lean, slender (descr., Latin), adj. ekari: Ekari language of New Guinea (other), noun app. elachistus, us, a, um: small (descr., Greek), adj. elatus, us, a, um: high (descr., Latin), adj. elburs: Elburs mountain range (geogr.), noun app. eldenbecki: D. Elden Beck (pers.), noun gen. elegans: elegant (descr., Latin), adj. un elegantula, us, a, um: neat (descr., Latin), adj. elenauerae: *Elenauera (pers.), noun gen. elevatus, us, a, um: elevated (descr., Latin), adj. elgae: *Elga Bamm (pers.), noun gen. elisabethae: *Elisabeth Hintelmann (pers.), noun gen. elizabethae: *Elizabeth (pers.), noun gen. eljasi: Mats Eljas (pers.), noun gen. ellai: *Ella Jean (pers.), noun gen. ellenbrookensis, is, is, e: Ellen Brook (geogr.), adj. name ellipticus, us, a, um: elliptical (descr., Greek), adj. ellypticus, us, a, um: elliptical (descr., Greek), adj. elmii: Mr. Karim Elmi (pers.), noun gen. elohri: Julius Flohr (not Elohr!), (pers.), noun gen. elongatulus, us, a, um: somewhat extended (descr., Latin), adj. elongatus, us, a, um: extended (descr., Latin), adj. elotus, us, a, um: elevated (elatus), (descr., Latin), adj. elusivus, us, a, um: rare (descr., Latin), adj. elutus, us, a, um: diluted (descr., Latin), adj. emaciatus, us, a, um: grown thin (descr., Latin), adj. emarginatus, us, a, um: notched (descr., Latin), adj. emilianus: us, a, um: Emilia (Emilia Romagna), (geogr.), adj. name eminens: eminent (descr., Latin), adj. emmae: *Emma Hendrich (wife of L. Hendrich), (pers.), noun gen. emmerichi: Emmerich Reitter (pers.), noun gen. endroedyi: Sebastian Endrödy-Younga (pers.), noun gen. enganensis, is, is, e: Enggano island (geogr.), adj. name englundi: Ronald A. Englund (pers.), noun gen. enigmaticus, us, a, um: enigmatic (descr., Latin), adj. enneagrammus, us, a, um: with 9 lines (descr., Greek), adj. epactus, us, a, um: added (descr., Greek), adj. ephippiiger: rider (descr., Greek), noun app. epibletus, us, a, um: imposing (descr., Greek), adj. epinephes: on clouds or mist (descr., Greek), adj. un epipleuricus, us, a, um: with epipleuron (descr., Greek), adj. erannus, us, a, um: lovely (descr., Greek), adj. eremita: hermit (descr., Greek), noun app. eretiformis, is, is, e: Eretes-like (descr.), adj. erichsoni/ -ii: Wilhelm Ferdinand Erichson (pers.), noun gen. ermaki: Ermak (Yermak Timofeyevich), (pers.), noun gen. ernsti: Ernst Wewalka (pers.), noun gen. errans: making error (descr., Latin), adj. erraticus, us, a, um: in error (descr., Latin), adj. erratus, us, a, um: in error (descr., Latin), adj. erteldi: Christian Erteld (pers.), noun gen. ertli: Johann Nepomuk Ertl (pers.), noun gen. eruditus, us, a, um: of richly red colour (descr., Greek), adj. erythraeus, us, a, um: Eritrea (geogr.), adj. name erythrocephalus, us, a, um: with head red (descr., Greek), adj. erythromelas: reddish black (descr., Greek), adj. un erythropus: red foot (descr., Greek), noun app. erythrostomus, us, a, um: with mouth red (descr., Greek), adj. erytropterus, us, a, um: with wings red (descr., Greek), adj. erzurumensis, is, is, e: Erzurum (geogr.), adj. name esau: Esau (hairy profemur), (descr.), noun app. escalerai: Manuel M. de Escalera (pers.), noun gen. escheri: Heinrich Escher-Zollikofer (pers.), noun gen. espanyoli: Francesco Español (pers.), noun gen. essoni: Esson Leon (pers.), noun gen. esteriensis, is, is, e: Cap Esterias (geogr.), adj. name estrellensis, is, is, e: Serra da Estrêla (geogr.), adj. name etnieri: David Etnier (pers.), noun gen. eucritus, us, a, um: readily chosen (descr., Greek), adj. eugeniae: *Eugenia Zaitzeva (first wife of F. Zaitzev), (pers.), noun gen. eugrammus, us, a, um: with well-developed lines (descr., Greek), adj. eupteryx: beautiful wings (descr., Greek), noun app. euryomus, us, a, um: with shoulder wide (descr., Greek), adj. eurypleuron: wide ribbed (broad epipleura), (descr., Greek), noun app. evanescens: fainting (descr., Latin), adj. evanidus, us, a, um: fainted (descr., Latin), adj. evelynae: *Evelyn Lunschien-Fery (wife of H. Fery), (pers.), noun gen. eversmanni: Eduard Friedrich von Eversmann (pers.), noun gen. exaratus, us, a, um: hollowed out (descr.), adj. excavatus, us, a, um: excavated (descr., Latin), adj. exclamationis: exclamation (descr., Latin), noun app. excoffieri: Joseph Claude Excoffier (pers.), noun gen. excrucians: harassment (descr., Latin), noun app. exiguus, us, a, um: small (descr., Latin), adj. exilis, is, is, e: thin, slender (descr., Latin), adj. eximius, us, a, um: selected (descr., Latin), adj. exochomoides: Exochomus-like (descr.), adj. un exoletus, us, a, um: obsolete (descr., Latin), adj. exornatus, us, a, um: ornamented (descr., Latin), adj. explanatus, us, a, um: widened (descr., Latin), adj. explicitus, us, a, um: disentangled (descr., Latin), adj. expositus, us, a, um: exposed (descr., Latin), adj. expressus, us, a, um: ready (descr., Latin), adj. exsanguis, is, is, e: anaemic, pale (descr., Latin), adj. exsoletus, us, a, um: obsolete (descr., Latin), adj. exspectatus, us, a, um: expected (descr., Latin), adj. extensus, us, a, um: extended (descr., Latin), adj. extenuans: underestimating (descr., Latin), adj. externepunctatus, us, a, um: externally punctate (descr., Latin), adj. externus, us, a, um: from abroad (descr., Latin), adj. extinctus, us, a, um: extinct (descr., Latin), adj. ezo: Ezo (Hokkaido), (geogr.), noun app. fabressei: Fabresse (pers.), noun gen. fabricii: Johann Christian Fabricius (pers.), noun gen. fabriciusi: Johann Christian Fabricius (pers.), noun gen. facetus, us, a, um: graceful (descr., Latin), adj. facilis, is, is, e: facile (easily come by), (descr., Latin), adj.

16 14 Anders N. Nilsson fairmairei: Léon Fairmaire (pers.), noun gen. falcozi: Louis Falcoz (pers.), noun gen. faldermanni: Franz Faldermann (pers.), noun gen. falkenstromi: Gustav Adolf Falkenström (pers.), noun gen. falklandicus, us, a, um: Falkland islands (geogr.), adj. name fallaciosus, us, a, um: deceitful (descr., Latin), adj. fallax: deceitful, (descr. Latin), adj. un falli: Henry Clinton Fall (pers.), noun gen. falsificus, us, a, um: false (descr., Latin), adj. fangensis, is, is, e: Fang (geogr.), adj. name farctus, us, a, um: well stuffed (descr., Latin), adj. faroensis, is, is, e: Faroes (geogr.), adj. name farquharensis, is, is, e: Farquhar atoll (geogr.), adj. name fartilis, is, is, e: fattened (descr., Latin), adj. fasciatocollis, is, is, e: with neck banded (descr., Latin), adj. fasciatus, us, a, um: banded (descr., Latin), adj. fascicollis, is, is, e: with neck banded (descr., Latin), adj. fasciventris, is, is, e: ventrally banded (descr., Latin), adj. fastidiosus, us, a, um: fastidious (descr., Latin), adj. fastuosus, us, a, um: splendid (descr., Latin), adj. fatigus, us, a, um: of much effort (descr., Latin), adj. fauconneti: Mary-Louis Fauconnet (pers.), noun gen. favareli: Geo Favarel (pers.), noun gen. feae: Leonardo Fea (pers.), noun gen. felixi: Félix Guignot (pers.), noun gen. femoralis, is, is, e: pertaining to femur (descr., Latin), adj. femoratus, us, a, um: pertaining to femur (descr., Latin), adj. fenestralis, is, is, e: with small window (transparent zone), (descr., Latin), adj. fenestratus, us, a, um: with small window (descr., Latin), adj. fengi: H. T. Feng (pers.), noun gen. fennicus, us, a, um: Fennia (Finland), (geogr.), adj. name feraudi: Féraud (pers.), noun gen. fernandensis, is, is, e: Fernando Po (geogr.), adj. name fernandoi: Fernando Pederzani (pers.), noun gen. ferrugatus, us, a, um: rust-coloured (descr., Latin), adj. ferrugineus, us, a, um: rust-coloured (descr., Latin), adj. ferruginicollis, is, is, e: with neck rust-coloured (descr., Latin), adj. ferus, us, a, um: severe (descr., Latin), adj. feryi: Hans Fery (pers.), noun gen. festae: Enrico Festa (pers.), noun gen. festivus, us, a, um: festive (descr., Latin), adj. fictor: one who shapes (descr., Latin), noun app. fidschiensis, is, is, e: Fidschi (Fiji), (geogr.), adj. name figuratus, us, a, um: figurative (descr., Latin), adj. fijiensis, is, is, e: Fiji (geogr.), adj. name filicornis, is, is, e: with antennae thread-like (descr., Latin), adj. filiformis, is, is, e: thread-like (descr., Latin), adj. filiolus: small thread (descr., Latin), noun app. fimbriatus, us, a, um: fringed (descr., Latin), adj. fimbriolatus, us, a, um: with fringes (descr., Latin), adj. finitimus, us, a, um: adjacent (descr., Latin), adj. finus: the last (sp. in revision), (descr.), noun app. fischeri: Ejnar Fischer (pers.), noun gen. [Megaporus] fischeri: Fischer (pers.), noun gen. [Hyphydrus] flammulatus, us, a, um: with small flames (descr., Latin), adj. flaveolus, us, a, um: yellowish (descr., Latin), adj. flavescens: becoming yellow (descr., Latin), adj. flavicans: being yellow (descr., Latin), adj. un flaviceps: with head yellow (descr., Latin), adj. un flavicollis, is, is, e: with neck yellow (descr., Latin), adj. flaviculus, us, a, um: small and yellow (descr., Latin), adj. flavidus, us, a, um: yellowish (descr., Latin), adj. flavipennis, is, is, e: with wings yellow (descr., Latin), adj. flavipes: yellow foot (descr., Latin), noun app. flaviventris, is, is, e: with belly yellow (descr., Latin), adj. flavobrunneus, us, a, um: yellowish brown (descr., Latin), adj. flavocinctus, us, a, um: with girdle yellow (descr., Latin), adj. flavofasciatus, us, a, um: yellow banded (descr., Latin), adj. flavogriseus, us, a, um: yellowish grey (descr., Latin), adj. flavoguttatus, us, a, um: yellow spotted (descr., Latin), adj. flavolineatus, is, is, e: yellow lined (descr., Latin), adj. flavomaculatus, us, a, um: with yellow markings (descr., Latin), adj. flavomarginatus, us, a, um: with yellow border (descr., Latin), adj. flavopictus, us, a, um: painted yellow (descr., Latin), adj. flavopunctatus, us, a, um: with yellow dots (descr., Latin), adj. flavoscriptus, us, a, um: with yellow markings (descr., Latin), adj. flavoscutatus, us, a, um: with yellow shield (descr., Latin), adj. flavoscutellatus, us, a, um: with small yellow shield (descr., Latin), adj. flavosignatus, us, a, um: with yellow mark (descr., Latin), adj. flavotestaceus, us, a, um: yellowish red (descr., Latin), adj. flavovittatus, us, a, um: with yellow stripe (descr., Latin), adj. flavus, us, a, um: yellow (descr., Latin), adj. flebilis, is, is, e: pitiful (descr., Latin), adj. fletcheri: James Fletcher (pers.), noun gen. flexuosus, us, a, um: with many bends (descr., Latin), adj. flindersii: Flinders (pers.), noun gen. flohri: Julius Flohr (pers.), noun gen. flohrianus, us, a, um: Julius Flohr (pers.), adj. name florencae: *Florence (pers.), noun gen. flores: Flores (geogr.), noun app. floridanus, us, a, um: Florida (geogr.), adj. name florissantensis, is, is, e: Florissant (geogr.), adj. name flumineus, us, a, um: pertaining to river (hab., Latin), adj. fluminum: of rivers (hab., Latin), noun gen. fluviaticus, us, a, um: in running water (hab., Latin), adj. fluviatilis, is, is, e: in running water (hab., Latin), adj. foetidus, us, a, um: mephitical, stinking (descr., Latin), adj. folkertsi: George W. Folkerts (pers.), noun gen. fontanus, us, a, um: from spring (hab., Latin), adj. fonticola: one that lives in spring (hab., Latin), noun app. fontinalis, is, is, e: from spring (hab., Latin), adj. fordii: Ford (pers.), noun gen. formaster, er, ra, rum: shaped in similar way (descr., Latin), adj. formosanus, us, a, um: Formosa (Taiwan), (geogr.), adj. name formosulus, us, a, um: small and beautiful (descr., Latin), adj. formosus, us, a, um: beautiful (descr., Latin), adj.

17 All diving beetle specific and subspecific names explained 15 forsteri: Johann Reinhold Forster (pers.), noun gen. forstromii: Forström (pers.), noun gen. fortis, is, is, e: strong (descr., Latin), adj. fortisspina: strong spines (descr., Latin), noun app. fortiterpunctatus, us, a, um: with strong punctation (descr., Latin), adj. fossarum: of ditches (descr., Latin), noun gen. fossiger, er, era, erum: from ditch (hab., Latin), adj. fossilis, is, is, e: fossile (descr., Latin), adj. fossulata, us, a, um: with small pits (descr., Latin), adj. fossulipennis, is, is, e: wing with small pits (descr., Latin), adj. fosteri: Garth N. Foster (pers.), noun gen. foveatus, us, a, um: with pits (descr., Latin), adj. foveiceps: head with pits (descr., Latin), adj. un foveola: small pit (descr., Latin), noun app. foveolatus, us, a, um: with small pits (descr., Latin), adj. fractifer, er, era, erum: carrying fracture (descr., Latin), adj. fractilinea: broken line (descr., Latin), noun app. fractistriatus, us, a, um: with fragmented scratches (descr., Latin), adj. fractivittis, is, is, e: with fragmented stripe (descr., Latin), adj. fractus, us, a, um: broken (descr., Latin), adj. fragilis, is, is, e: fragile (descr., Latin), adj. fragrans: fragrant (descr., Latin), adj. francescae: *Frances Young (pers.), noun gen. franki: Frank N. Young Jr. (pers.), noun gen. franklinii: John Franklin (pers.), noun gen. franzi: Herbert Franz (pers.), noun gen. frater: brother (descr., Latin), noun app. fraterculus: dear brother (descr., Latin), noun app. fraternus, us, a, um: like brothers (descr., Latin), adj. fraudator: impostor (descr., Latin), noun app. fresnedai: Javier Fresneda (pers.), noun gen. freudei: Heinz Freude (pers.), noun gen. freyi: G. Frey (pers.), noun gen. fridaywellensis, is, is, e: Friday well (geogr.), adj. name friedrichi: W. Friedrich (pers.), noun gen. [Gaurodytes, Rhantus] friedrichi: Friedrich Wewalka (pers.), noun gen. [Allopachria] frigidus, us, a, um: cold (descr., Latin), adj. frischii: Johann Leonard Frisch (pers.), noun gen. frisius, us, a, um: Frisian islands (geogr.), adj. name froehlichi: Fritz Fröhlich (pers.), noun gen. frontalis, is, is, e: relating to forehead (descr., Latin), adj. frustrator: impostor (descr., Latin), noun app. fryi/ -ii: Alexander Fry (pers.), noun gen. fufai: Fufa (boy), (pers.), noun gen. fulgidus, us, a, um: shining (descr., Latin), adj. fuliginosus, us, a, um: very sooty (descr., Latin), adj. fulvaster, er, ra, rum: red brownish (descr., Latin), adj. fulvescens: becoming reddish brown (descr., Latin), adj. fulviceps: with head yellowish-brown (descr.), adj. un fulvicollis, is, is, e: with neck yellowish-brown (descr., Latin), adj. fulvipennis, is, is, e: with wings yellowish-brown (descr., Latin), adj. fulviventris, is, is, e: with belly yellowish-brown (descr., Latin), adj. fulvoguttatus, us, a, um: with yellowish-brown spots (descr., Latin), adj. fulvonotatus, us, a, um: dorsally yellowish-brown (descr., Latin), adj. fulvosparsus, us, a, um: yellowish-brown and relatively rare (descr., Latin), adj. fulvus, us, a, um: yellowish-brown (descr., Latin), adj. fumatus, us, a, um: smoke-coloured (descr., Latin), adj. fume: spring (fu-me), (hab.), noun app. funebris, is, is, e: fatal (descr., Latin), adj. funereus, us, a, um: fatal (descr., Latin), adj. funestus, us, a, um: fatal (descr., Latin), adj. funkii: Michael Funk (pers.), noun gen. furvus, us, a, um: dark (descr., Latin), adj. fuscatus, us, a, um: darkened (descr., Latin), adj. fuscicollis, is, is, e: with neck dark (descr., Latin), adj. fuscidorsis, is, is, e: with back dark (descr., Latin), adj. fuscinus, us, a, um: darkish (descr., Latin), adj. fuscipennis, is, is, e: with wings dark (descr., Latin), adj. fuscipes: black foot (descr., Latin), noun app. fuscitarsis, is, is, e: with foot black (descr., Latin), adj. fusciventris, is, is, e: with belly dark (descr., Latin), adj. fuscoaenescens: becoming dark copper-tinged (descr., Latin), adj. fuscobrunneus, us, a, um: dark brown (descr., Latin), adj. fuscolineatus, us, a, um: with dark lines (descr., Latin), adj. fuscomaculatus, us, a, um: with dark markings (descr., Latin), adj. fuscorufus, us, a, um: dark red (descr., Latin), adj. fuscostriatus, us, a, um: with dark scratches (descr., Latin), adj. fuscotestaceus, us, a, um: dark orange red (descr., Latin), adj. fusculus, us, a, um: small and dark (descr., Latin), adj. fuscus, us, a, um: dark-coloured (descr., Latin), adj. gabonicus, us, a, um: Gabon (geogr.), adj. name gabrielae: *Gabriela Gabi Seiler (pers.), noun gen. gagates: vitrain (descr., Latin), noun app. gagatinus, us, a, um: like vitrain, jet black (descr., Latin), adj. galamensis, is, is, e: Galama mountains (geogr.), adj. name galapagoensis, is, is, e: Galapagos islands (geogr.), adj. name galbinatus, us, a, um: made greenish yellow (descr., Latin), adj. galileae: Galilee (geogr.), noun gen. galla: Galla people of Ethiopia (people), noun app. gallicus, us, a, um: Gallia (geogr.), adj. name galpini: Mr. Galpin (pers.), noun gen. ganglbaueri: Ludwig Ganglbauer (pers.), noun gen. gapa: Cunninghams Gap (geogr.), noun app. garambanus, us, a, um: Garamba NP (geogr.), adj. name gardineri: J. Stanley Gardiner (pers.), noun gen. gardnerii: John Gardner (pers.), noun gen. gaudens: pleasing (descr., Latin), adj. gaudichaudii: Gaudichaud (pers.), noun gen. gayndahensis, is, is, e: Gayndah (geogr.), adj. name geayi: F. Geay (pers.), noun gen. geetae: *Geet (pers.), Indian, noun gen. geijskesi: Dirk Cornelius Geijskes (pers.), noun gen. geiseri: Michael Geiseri (pers.), noun gen. gelidus, us, a, um: cold (hab., Latin), adj. gemellatus, us, a, um: doubled (descr., Latin), adj.

18 16 Anders N. Nilsson gemellus, us, a, um: like small pearl (descr., Latin), adj. geminatus, us, a, um: doubled (descr., Latin), adj. geminodes: twin-like (descr., Latin), adj. geminus: twin (descr., Latin), noun app. genei: Carlo Guiseppe Gené (pers.), noun gen. geniculatus, us, a, um: with small knots (descr., Latin), adj. gentilis, is, is, e: noble (descr., Latin), adj. georgicus, us, a, um: Georgia (nation), (geogr.), adj. name georginae: *Georgina Watts (pers.), noun gen. gereckei: Reinhard Gerecke (collector), (pers.), noun gen. germanus, us, a, um: authentic, closely related (descr., Latin), adj. gestroi: Raphael Gestro (pers.), noun gen. ghanensis, is, is, e: Ghana (geogr.), adj. name ghesquierei: J. Ghesquière (pers.), noun gen. gibbi: *Geraldine Barter (G.B.), (pers.), noun gen. gibbicollis, is, is, e: with neck vaulted (descr., Latin), adj. gibbosus, us, a, um: humped (descr., Latin), adj. gibbulus, us, a, um: small and vaulted (descr., Latin), adj. gibbus, us, a, um: vaulted (descr., Latin), adj. gibsoni: K.E. Gibson (pers.), noun gen. giganteus, us, a, um: gigantic (descr., Greek), adj. gigantoides: giganteus-like (descr.), adj. un gigas: giant (descr., Latin), noun app. gignouxi: Gignoux family (pers.), noun gen. gignouxorum: Gignoux family (pers.), noun gen. gilbertii: John Gilbert (pers.), noun gen. gilvus, us, a, um: pale yellow (descr., Latin), adj. girardi: Cl. Girard (pers.), noun gen. gittidasi: Gittidas (geogr.), noun gen. giuliani: D.D. Guiliani (pers.), noun app. glabellus, us, a, um: smooth (descr., Latin), adj. glaber, er, ra, rum: bald (descr., Latin), adj. glabratus, us, a, um: made bald (descr., Latin), adj. glabrellus, us, a, um: somewhat bald (descr., Latin), adj. glabrescens: becoming bald (descr., Latin), adj. glabricollis, is, is, e: with neck bald (descr., Latin), adj. glabricula, us, a, um: somewhat bald (descr., Latin), adj. glabriusculus, us, a, um: almost bald (descr., Latin), adj. glacialis, is, is, e: glacial (hab., Latin), adj. glasunovi: Glazunov (pers.), noun gen. glaucus, us, a, um: bluish green (descr., Latin), adj. glazunovi: Glazunov (pers.), noun gen. globosus, us, a, um: globe-like (descr., Latin), adj. globulosus, us, a, um: globe-like (descr., Latin), adj. glomeratus, us, a, um: coiled up (descr., Latin), adj. glyphicus, us, a, um: chiseled (descr., Greek), adj. glypta, us, a, um: engraved (descr., Greek), adj. godeffroyi: J.P. Godeffroy (pers.), noun gen. godmanni: Godman (pers.), noun gen. goedelii: Gödel (pers.), noun gen. gogonensis, is, is, e: Sha Gogona (geogr.), adj. name goias: Goiás (geogr.), noun app. goldbergi: John Goldberg (pers.), noun gen. goldschmidti: Richard Goldschmidt (pers.), noun gen. [Hydroporus] goldschmidti: Tom Goldschmidt (pers.), noun gen. [Herophydrus] gomerensis, is, is, e: La Gomera (geogr.), adj. name gondaricus, us, a, um: Gondar (geogr.), adj. name gory: Hippolyte Louis Gory (pers.), noun app. gossei: Philip Henry Gosse (pers.), noun gen. gotschii: Gotsch (pers.), noun gen. gottingensis, is, is, e: Göttingen (geogr.), adj. name gottwaldi: Stephan Gottwald (pers.), noun gen. goudoti/ -ii: Justin Goudot (pers.), noun gen. gougeletii: Jean Scipio Gougelet (pers.), noun gen. gouini: F. Gouin (pers.), noun gen. gounellei: Pierre Emile Gounelle (pers.), noun gen. grabowskyi: Fritz Grabowsky (Grabowski), (pers.), noun gen. gracilicornis, is, is, e: with antennae slender (descr., Latin), adj. gracilis, is, is, e: slender (descr., Latin), adj. graellsi: Mariano de la Paz Graëlls (pers.), noun gen. gramineus, us, a, um: like grass (descr., Latin), adj. grammicus, us, a, um: with lines (descr., Greek), adj. grammopterus, us, a, um: with lines on wings (descr., Greek), adj. granarius, us, a, um: seed-like (descr., Latin), adj. grandicollis, is, is, e: with neck large (descr., Latin), adj. grandis, is, is, e: large (descr., Latin), adj. grandisinuatus, us, a, um: large and S-shaped (descr., Latin), adj. granoides: grana-like (descr.), adj. un granosus, us, a, um: full of seeds (descr., Latin), adj. granularis, is, is, e: with small grains (descr., Latin), adj. granulatus, us, a, um: with small grains (descr., Latin), adj. granulum: small seed (descr., Latin), noun app. granum: seed (descr., Latin), noun app. grapei: Zacharias Grape (pers.), noun gen. grapii: Zacharias Grape (pers.), noun gen. graueri: Rudolf Grauer (pers.), noun gen. gravicornis, is, is, e: with antennae heavy (descr., Latin), adj. gravidus, us, a, um: stuffed (descr., Latin), adj. gravis, is, is, e: heavy (descr., Latin), adj. gredensis, is, is, e: Sierra de Gredos (geogr.), adj. name gregarius, us, a, um: clustered (descr., Latin), adj. gridellii: Edoardo Gridelli (pers.), noun gen. griffinii: Achille Griffini (pers.), noun gen. gringo: foreigner (other, Spanish), noun app. griphodes: like a riddle (descr., Latin), adj. griseipennis, is, is, e: with wings grey (descr., Latin), adj. griseoguttatus, us, a, um: with grey spots (descr., Latin), adj. griseostriatus, us, a, um: with grey scratches (descr., Latin), adj. griseus, us, a, um: grey (descr., Latin), adj. groenlandicus, us, a, um: Grönland (Greenland), (geogr.), adj. name gronovii: Laurentius Theodorus Gronovius (pers.), noun gen. grossepunctatus, us, a, um: with coarse punctation (descr., Latin), adj. grossula, us, a, um: rather big (descr., Latin), adj. grossus, us, a, um: big (descr., Latin), adj. grouvellei: Antoine Grouvelle (pers.), noun gen. gschwendtneri: Ludwig Gschwendtner (pers.), noun gen. guadalcanalensis, is, is, e: Guadalcanal (geogr.), adj. name guadelupensis, is, is, e: Guadelupe (geogr.), adj. name

19 All diving beetle specific and subspecific names explained 17 guatemalensis, is, is, e: Guatemala (geogr.), adj. name guayanensis, is, is, e: Guayana (geogr.), adj. name guayaquilensis, is, is, e: Guayaquil (geogr.), adj. name gueorguievi: Vassil Guéorguiev (pers.), noun gen. guerini: Félix Édouard Guérin-Méneville (pers.), noun gen. guernei: J. de Guerne (pers.), noun gen. guerryi: Paul Guerry (pers.), noun gen. guidettii: Walter Guidetti (pers.), noun gen. guidi: Guy (grandchild), (pers.), noun gen. guignoti: Félix Guignot (pers.), noun gen. guignotianus, us, a, um: Félix Guignot (pers.), adj. name guineensis, is, is, e: Guinea (geogr.), adj. name guizhouensis, is, is, e: Guizhou (geogr.), adj. name gujaratensis, is, is, e: Gujarat (geogr.), adj. name gumwellensis, is, is, e: Gum well (geogr.), adj. name gusenleitneri: F. Gusenleitner (pers.), noun gen. gutta: drop (drop-like spot), (descr., Latin), noun app. guttalis, is, is, e: spotted (descr., Latin), adj. guttatus, us, a, um: spotted (descr., Latin), adj. gutteridgei: Rob Gutteridge (pers.), noun gen. gutticollis, is, is, e: with neck spotted (descr., Latin), adj. guttiger, er, era, erum: carrying drop (descr., Latin), adj. guttulatus, us, a, um: with small drop (descr., Latin), adj. guttulosus, us, a, um: with small drop (descr., Latin), adj. guttulus, us, a, um: with small drop (descr., Latin), adj. gyllenhalii: Leonhard Gyllenhal (pers.), noun gen. gyrinoides: Gyrinus-like (descr.), adj. un haagi: Georg Johann Haag (pers.), noun gen. habelmanni: Paul Sigismund Habelmann (pers.), noun gen. habilis, is, is, e: apt, fit (descr., Latin), adj. hades: god of the dead (myth., Greek), noun app. haeffneri: Johann Christian Friedrich Haeffner (pers.), noun gen. haemorrhoidalis, is, is, e: blood-coloured in river (descr., Greek), adj. hafti: Jan Haft (pers.), noun gen. hahni: Hans Jurgen Hahn (pers.), noun gen. hainanensis, is, is, e: Hainan (geogr.), adj. name hajeki: Jiří Hájek (pers.), noun gen. hakkariensis, is, is, e: Hakkari province (geogr.), adj. name halensis, is, is, e: Hale (Halle), (geogr.), adj. name hamadae: Neusa Hamada (pers.), noun gen. hamatus, us, a, um: barbed (descr., Latin), adj. hamifer, er, era, erum: carrying hook (descr., Latin), adj. hamoni: Harold Hamon (pers.), noun gen. hamulatus, us, a, um: with small hooks (descr., Latin), adj. hancocki: G.L.R. Hancock (pers.), noun gen. hansardii: Hansard (pers.), noun gen. haraldi: Harald Lindberg (pers.), noun gen. hararensis, is, is, e: Harare (geogr.), adj. name hardenbergi: C.B. Hardenberg (pers.), noun gen. hardyi: Mr. Hardy (pers.), noun gen. harleyi: Harley Barron (pers.), noun gen. haroldi: Harold M. Miller (pers.), noun gen. harrarensis, is, is, e: Harrare (geogr.), adj. name harrisii: Thaddeus William Harris (pers.), noun gen. harrisoni: Mr. A.D. Harrison (pers.), noun gen. hasegawai: Michiaki Hasegawa (pers.), noun gen. hastatus, us, a, um: armed with spear (descr., Latin), adj. hatchi: Melville Hatch (pers.), noun gen. hautmanni: *Susanne & Andreas Hautmann (pers.), noun gen. hautmannorum: *Susanne & Andreas Hautmann (pers.), noun gen. havaniensis, is, is, e: Havana (geogr.), adj. name hebaueri: Franz & Hans Hebauer (pers.), noun gen. [Deronectes] hebaueri: Hans Hebauer (pers.), noun gen. [Hydroporus] hebauerorum: Franz & Hans Hebauer (pers.), noun gen. hebes: blunt (descr., Latin), adj. un hebeter: the blunt one (descr., Latin), noun app. hebraicus, us, a, um: jewish (people, Latin), adj. hedini: Sven Hedin (pers.), noun gen. hedwigae: *Hedwig Gruber (pers.), noun gen. hedwigi: *Hedwig Gruber (pers.), noun gen. hedychrous, us, a, um: sweet-smelling (descr., Latin), adj. heeri: Oswald Heer (pers.), noun gen. heggiensis, is, is, e: Heggie s Rock (geogr.), adj. name heidiae: *Heidi Brancucci (pers.), noun gen. [Laccophilus] heidiae: *Heidi Riens (pers.), noun gen. [Copelatus] heinertzi: R. Heinertz (pers.), noun gen. hellenicus, us, a, um: Hellas (Greece), (geogr.), adj. name helocrinus, us, a, um: from helocrene spring (hab.), adj. helveticus, us, a, um: Helvetia (Switzerland), (geogr.), adj. name helvolus, us, a, um: pale yellow green (descr., Latin), adj. hemani: Heman (pers., Indian), noun gen. hendrichi: Lars Hendrich (pers.), noun gen. herbeti: Herbet (pers.), noun gen. hermanni: Joh. Hermann (pers.), noun gen. heros: hero (descr., Greek), noun app. herrmanni: Joh. Herrmann (pers.), noun gen. heterocerus, us, a, um: with antennae different (descr., Greek), adj. heterogynus, us, a, um: with female different (descr., Greek), adj. heteronychus, us, a, um: with claws different (descr.), adj. heubergeri: Max Heuberger (pers.), noun gen. heydeni: Lucas von Heyden (pers.), noun gen. hiekei: Fritz Hieke (pers.), noun gen. hieroglyphicus, us, a, um: with hieroglyphs (descr., Greek), adj. hillviewensis, is, is, e: Hillview station (geogr.), adj. name hinkleri: Hinkler Well catchment (geogr.), noun gen. hintelmannae: *Elisabeth Hintelmann (pers.), noun gen. hintoni: Howard Everett Hinton (pers.), noun gen. hinzeae: *Susan Hinzae (pers.), noun gen. hirsutus, us, a, um: with distinct pubescence (descr., Latin), adj. hirtellus, us, a, um: somewhat shaggy (descr., Latin), adj. hirtipes: shaggy foot (descr., Latin), noun app. hisamatsui: S. Hisamatsu (pers.), noun gen. hispanicus, us, a, um: Hispania (Spain), (geogr.), adj. name histeroides: Hister-like (descr.), adj. un histrio: actor (descr., Latin), noun app. hiurai: I. Hiura (pers.), noun gen. hobbsi: Horton H. Hobbs Jr. (pers.), noun gen. hochhuti: Johann Heinrich Hochhut (pers.), noun gen.

20 18 Anders N. Nilsson hochi: Karl Hoch (pers.), noun gen. hockingi: Brian Hocking (pers.), noun gen. hokkaidensis, is, is, e: Hokkaido (geogr.), adj. name hollingsworthi: Rod Hollingsworth (pers.), noun gen. holmeni: Mogens Holmen (pers.), noun gen. holomelas: completely black (descr., Greek), adj. un holosericeus, us, a, um: from pure silk (descr., Greek), adj. holzmanni: *Chantale & Robert Holzmann (pers.), noun gen. holzmannorum: *Chantale & Robert Holzmann (pers.), noun gen. holzschuhi: Carolus Holzschuh (pers.), noun gen. homoenyx: uniform claws (descr., Greek), noun app. hookeri: Joseph Dalton Hooker (pers.), noun gen. hopffgarteni: Baron Georg Max v. Hopffgarten (pers.), noun gen. hoppingi: Ralph Hopping (pers.), noun gen. horai: S.L. Hora (pers.), noun gen. horni: George Henry Horn (pers.), noun gen. [Laccophilus Branden] horni: Walther Hermann Richard Horn (pers.), noun gen. [Copelatus, Laccophilus Rég.] hornii: George Henry Horn (pers.), noun gen. hortense: *Hortense Griffin Lanphere (pers.), noun app. hostilis, is, is, e: foreign (descr., Latin), adj. hottentottus, us, a, um: Khoikhoi people of Cape (people), adj. hova: Hova or Merina people of Madagascar (people), noun app. howittii: Alfred William Howitt (pers.), noun gen. hozgargantae: Hozgarganta river (geogr.), noun gen. huastecus, us, a, um: Huastec people of Mexico (people), adj. name hubbardi: David A. Hubbard (pers.), noun gen. hubbelli: Theodore H. Hubbell (pers.), noun gen. hudsonicus, us, a, um: Hudson Bay (geogr.), adj. name huehni: Hühn (pers.), noun gen. hulae: Hula reserve (geogr.), noun gen. humboldtensis, is, is, e: Humboldt county (geogr.), adj. name humeralis, is, is, e: pertaining to shoulder (descr., Latin), adj. humilis, is, is, e: low (descr., Latin), adj. hummeli: Arvid David Hummel (pers.), noun gen. hunteri: William Hunter, (pers.) noun gen. huttoni: Prof. Hutton (pers.), noun gen. hyalinus, us, a, um: transparent, like glass (greenish), (descr., Greek), adj. hybneri: Johann Gottfried Hübner (pers.), noun gen. hybridus, us, a, um: of mixed race (descr., Latin), adj. hydaticoides: Hydaticus-like (descr.), adj. un hydrophiloides: Hydrophilus-like (descr.), adj. un hydropicus, us, a, um: visible in water (descr., Latin), adj. hydroporoides: Hydroporus-like (descr.), adj. un hydrovatoides: Hydrovatus-like (descr.), adj. un hygrobioides: Hygrobia-like (descr.), adj. un hygrobius, us, a, um: living in water (hab., Latin), adj. hygrotoides: Hygrotus-like (descr.), adj. un hylaeus, us, a, um: living in forest (hab., Greek), adj. hylobates: deep forest (hab., Greek), noun app. hynesi: H.B. Noel Hynes (pers.), noun gen. hyperboreus, us, a, um: northern (hab., Latin), adj. hyphoporoides: Hyphoporus-like (descr.), adj. un hyphydroides: Hyphydrus-like (descr.), adj. un hypochlorus, us, a, um: ventrally green (descr., Greek), adj. hypomelas: ventrally black (descr., Greek), adj. un ibaguensis, is, is, e: Ibagué (geogr.), adj. name ibarrai: Jorge Ibarra (pers.), noun gen. ibericus, us, a, um: Iberica (geogr.), adj. name ibonum: Ibo people of Nigeria (people), noun gen. ibrahimi: Ibrahim Mohamed Iman (pers.), noun gen. idanus, us, a, um: fair, lovely (descr., Greek), adj. ifnii: Ifni lake (geogr.), noun gen. ifranensis, is, is, e: Ifrane (geogr.), adj. name ignarus, us, a, um: red like fire (descr., Latin), adj. ignobilis, is, is, e: unknown (descr., Latin), adj. ignoratus, us, a, um: ignored (descr., Latin), adj. ignotus, us, a, um: unknown (descr., Latin), adj. iguelaensis, is, is, e: Iguéla lagoon (geogr.), adj. name ijimai: Mr. K. Ijima (pers.), noun gen. ikedai: Mr. T. Ikeda (pers.), noun gen. illigeri: Johann Karl Wilhelm Illiger (pers.), noun gen. ilybiiforme, is, is, e: shaped like an Ilybius (descr.), adj. ilybioides: Ilybius-like (descr., Latin), adj. un imasakai: S. Imasaka (pers.), noun gen. imbricata, us, a, um: tiled (descr., Latin), adj. imitator: imitator (descr., Latin), noun app. imitatrix: female imitator (descr., Latin), noun app. immaculatus, us, a, um: without marks (descr., Latin), adj. immaculipennis, us, a, um: wings without marks (descr., Latin), adj. immarginatus, us, a, um: without border (descr., Latin), adj. immaturus, us, a, um: immature (descr., Latin), adj. immitis, is, is, e: not soft (descr., Latin), adj. immundus, us, a, um: dirty (descr., Latin), adj. immunis, is, is, e: free (descr., Latin), adj. impar: not similar (descr., Latin), adj. un imperfectus, us, a, um: immature (descr., Latin), adj. imperialis, is, is, e: magnificent (descr., Latin), adj. impressicollis, is, is, e: with neck impressed (descr., Latin), adj. impressifrons: impressed frons (descr., Latin), noun app. impressopunctatus, us, a, um: with impressed punctation (descr., Latin), adj. impressus, us, a, um: impressed (descr., Latin), adj. impunctatus, us, a, um: without punctation (descr., Latin), adj. inaciculatus, us, a, um: not scratched with needle (descr., Latin), adj. inaequalis, is, is, e: unlike (descr., Latin), adj. inagua: Great Inagua island (geogr.), noun app. inanimatus, us, a, um: killed (descr., Latin), adj. incarinatus, us, a, um: not ridged (descr., Latin), adj. incaustus, us, a, um: not burnt (descr., Greek), adj. incertus, us, a, um: uncertain (descr., Latin), adj. incisus, us, a, um: carved (descr., Latin), adj. includens: becoming included (descr., Latin), adj. incognitus, us, a, um: unknown (descr., Latin), adj. incommodus, us, a, um: uncomfortable (caused much work), (descr., Latin), adj. incompleta, us, a, um: not complete (descr., Latin), adj. inconspectus, us, a, um: inconsiderate (descr., Latin), adj. inconspicuus, us, a, um: insignificant (descr., Latin), adj.

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