1 click for previous page Perciformes: Percoidei: Cirrhitidae 3321 CIRRHITIDAE Hawkfishes by J.E. Randall Diagnostic characters: Oblong fishes (size to about 30 cm), body depth 2 to 4.6 times in standard length. A fringe of cirri on posterior edge of anterior nostril. Two indistinct spines on opercle. A row of canine teeth in jaws, the longest usually anteriorly in upper jaw and half-way back on lower jaw; a band of villiform teeth inside the canines, broader anteriorly (in lower jaw only anteriorly). One or more cirri projecting from tips of interspinous membranes of dorsal fin. Dorsal fin continuous, with X spines and 11 to 17 soft rays, notched between spinous and soft portions; anal fin with III spines and 5 to 7 (usually 6) soft rays; pectoral fins with 14 rays, the lower 5 to 7 rays unbranched and usually enlarged, with the membranes deeply incised; pelvic fins with I spine and 5 soft rays. Principal caudal-fin rays 15. Branchiostegal rays 6. Scales cycloid. Swimbladder absent. Vertebrae 26. Colour: variable with species. cirri lower pectoral-fin rays thickened and unbranched Remarks: The hawkfish family consists of 10 genera and 38 species, 33 of which occur in the Indo-Pacific region; 19 species are found in the Western Central Pacific. Habitat, biology, and fisheries: Cirrhitids are bottom-dwelling fishes of coral reefs or rocky substrata; the majority occur in shallow water. They use their thickened lower pectoral-fin rays to wedge themselves in position in areas subject to surge. All species are carnivorous, feeding mainly on benthic crustaceans. Cyprinocirrhitus polyactis, however, is primarily a zooplankton-feeder (though still often seen at rest on the bottom). Most hawkfishes are small; separate species accounts are given below for the 3 largest hawkfishes that are occasionally used as food. Some of the smaller species, notably the bright red Neocirrhites armatus and the long-snouted Oxycirrhites typus, are of value as aquarium fishes. Similar families Cirrhitidae occurring in the area Serranidae: lower 5 to 7 pectoral-fin rays not simple and thickened (except for a few Plectranthias); no fringe of cirri on posterior nostril; scales usually ctenoid; usually III spines on opercle; vertebrae typically 24; swimbladder present. Serranidae (Plectranthias)
2 3322 Bony Fishes Key to the genera of Cirrhitidae occurring in the area 1a. Snout elongate, its length about 2 times in head length (Fig. 1); body slender, its depth 4.4 to 4.6 times in standard length; canine teeth in jaws only slightly longer than inner villiform teeth and nearly uniform in size Oxycirrhites 1b. Snout not elongate, its length 2.8 to 5 times in head length; body not slender, its depth 2 to 3.4 times in standard length; canine teeth in jaws markedly longer than inner villiform teeth, those at front of upper jaw and side of lower jaw enlarged a. Caudal fin lunate, the lobes produced (Fig. 2); soft dorsal-fin rays 16 or Cyprinocirrhites 2b. Caudal fin rounded, truncate, or slightly emarginate; soft dorsal-fin rays 11 to Fig. 1 Oxycirrhites Fig. 2 Cyprinocirrhites 3a. No large scales on cheek (small scales in more than 12 irregular rows) (Fig. 3) b. Four to 6 rows of large scales on cheek (small scales usually present as well) (Fig. 4)... 5 Fig. 3 Fig. 4 4a. Body not deep and not very compressed, its depth 2.6 to 3.4 times in standard length; body width 1.5 to 1.9 times in body depth; soft dorsal-fin rays 11 (rarely 12); palatine teeth present; upper margin of preopercle finely serrate or smooth; lower 7 pectoral-fin rays unbranched Cirrhitus 4b. Body deep and compressed, its depth 2 to 2.4 times in standard length; body width 2.9 to 3.1 times in body depth; soft dorsal-fin rays 13; palatine teeth absent; upper margin of preopercle coarsely serrate; lower 6 pectoral-fin rays unbranched Neocirrhites 5a. Five rows of large scales between lateral line and spinous portion of dorsal fin; a single cirrus from membrane near tip of each dorsal-fin spine (Fig. 5a) membranes of spinous portion of dorsal fin not deeply incised, those between longest dorsal-fi n spines extending 4/5 or more of distance from base to tip of spines; palatine teeth absent Paracirrhites 5b. Three or 4 rows of large scales between lateral line and base of spinous portion of dorsal fin; a tuft of cirri from membrane near tip of each dorsal-fin spine (Fig. 5b); membranes of spinous portion of dorsal fin deeply incised, those between longest spines less than 2/3 distance from base to tip of spines; palatine teeth present or absent... 6 a) Paracirrhites b) Cirrhitops Fig. 5 spinous part of dorsal fin
3 Perciformes: Percoidei: Cirrhitidae a. Soft dorsal-fin rays 14 (rarely 15); first 2 pectoral-fin rays unbranched; preorbital with a free posterior margin; interorbital not scaly; snout not pointed, the profile from interorbital to upper lip convex Cirrhitops 6b. Soft dorsal-fin rays 11 or 12 (rarely 13); first pectoral-fin ray unbranched, the second usually branched; preorbital with or without a free posterior margin; interorbital scaly or not scaly; snout pointed or not pointed a. Palatine teeth absent; longest dorsal-fin spine 3.5 to 4 times in body depth; snout not pointed Isocirrhitus 7b. Palatine teeth present; longest dorsal-fin spine 1.7 to 3.2 times in body depth; snout pointed a. Preopercular margin finely serrate (Fig. 6); preorbital without a free hind margin; interorbital scaly; first soft dorsal-fin ray not produced to a filament; lower 5 (rarely 6) pectoralfin rays unbranched Amblycirrhitus 8b. Preopercular margin coarsely serrate (Fig. 7); preorbital with hind margin free for about 1/4 to 1/2 distance from its lower edge to eye; interorbital not scaly; first soft dorsal-fin ray usually produced into a filament; lower 6 or 7 pectoral-fin rays unbranched.... Cirrhitichthys coarsely serrate Fig. 6 Amblycirrhitus Fig. 7 Cirrhitichthys Key to the species of Amblycirrhitus occurring in the area 1a. Lateral-line scales 48 to 50; tips of membranes of spinous portion of dorsal fin black Amblycirrhitus unimaculata (Ryukyu Islands, Taiwan Province of China, and Samoa) 1b. Lateral-line scales 39 to 42; tips of membranes of spinous portion of dorsal fin not black a. A large ocellated black spot on opercle; head not crossed by 3 dark bars.. Amblycirrhitus bimacula (Indo-Pacific) 2b. No large black spot on opercle; head crossed by 3 dark bars, the first passing ventrally from eye, the next 2 on operculum Amblycirrhitus oxyrhynchos (Goram Islands, Indonesia) Key to the species of Cirrhitichthys occurring in the area 1a. Body depth 2.4 to 2.7 times in standard length; bony interorbital space not narrow, its least width in adults about 1.7 times in eye diameter; free edge of preorbital usually with 1 to a few small spines; large dark spots on body, often coalescing to form 6 irregular bars; a pale-edged black spot larger than pupil on opercle at level of eye; 5 or 6 narrow oblique to diagonal dark bands on lower half of head, 3 or 4 of which extend ventroanteriorly from eye Cirrhitichthys aprinus (southern Japan to Great Barrier Reef) 1b. Body depth 2.7 to 3 times in standard length; bony interorbital space narrow, its least width about 2 times in eye diameter; free hind edge of preorbital without spines; colour not as above... 2
4 3324 Bony Fishes 2a. Rear edge of maxilla slightly anterior to a vertical at front edge of orbit; fourth dorsal-fin spine longest; first soft dorsal-fin ray not decidedly longer than other rays; body with numerous small red to reddish brown spots, many of which join to form about 7 bars; first bar vertical, centred on origin of dorsal fin and containing dark brown spots in its upper half, the other bars more oblique; 2 red to reddish brown bands extending ventroanteriorly from eye and a median band on snout Cirrhitichthys falco (western Pacific to Maldive Islands) 2b. Rear edge of maxilla reaching to or beyond a vertical at front edge of orbit; fifth dorsal-fin spine longest; first soft dorsal-fin ray decidedly longer than more posterior rays (3.5 cm standard length or larger); body with large close-set red to reddish brown blotches in 4 longitudinal rows and an additional row of smaller blotches along lateral line; head with numerous small red to reddish brown spots Cirrhitichthys oxycephalus (Indo-Pacific and tropical eastern Pacific) Key to the species of Cirrhitus occurring in the area 1a. Supraoccipital crest visible as a low ridge; no scales on interorbital space; no small white spots on head, body, and fins (white blotches nearly as large as eye present on body) Cirrhitus pinnulatus (Indo-Pacific) 1b. Supraorbital crest not visible externally; a narrow median band of scales on interorbital space; small white spots on head, body, and fins, those on body arranged in about 12 longitudinal rows Cirrhitus albopunctatus (Niuafoo Island, near Tonga Islands) Key to the species of Paracirrhites occurring in the area 1a. Second pectoral-fin ray branched, at least near tip; scales present on snout anterior to nostrils; no colour mark extending diagonally upward from rear part of eye b. Second pectoral-fin ray unbranched; no scales on snout anterior to nostrils; a prominent elongate solid or U-shaped dark mark extending diagonally upward from rear part of eye a. Tenth dorsal-fin spine slightly longer than ninth; lateral-line scales 45 to 49; numerous small dark spots on head and anteriorly on body Paracirrhites forsteri (Indo-Pacific) 2b. Tenth dorsal-fin spine notably longer than ninth; lateral-line scales 48 to 51; numerous small dark spots on body but none on head Paracirrhites hemistictus (Oceania except Hawaii; Great Barrier Reef, Christmas Island, and Cocos-Keeling Islands) 3a. Three dark-edged pale (orange in life) transverse bands on interopercle; postocular mark not black within U-shaped border (though may be darker than rest of head); 1 colour form with a broad pale stripe, lacking a black border, posteriorly on upper side; lateral-line scales 45 to Paracirrhites arcatus (Indo-Pacific) 3b. No transverse bands on interopercle; postocular mark black within its U-shaped border; a broad pale stripe, if present on upper side, broadly bordered with black; lateral-line scales 48 to a. Uniformly pale (bright yellow in life); black postocular mark narrower than pupil, originating at or below centre of rear edge of eye, its lower border not extending anterior to eye; no dark-edged white spots anteriorly on snout; no markings on maxilla; no white spot at anterior nostril Paracirrhites xanthus (Tuamotu Archipelago, Society Islands, and Marquesas) 4b. Light to dark brown (not yellow in life); postocular mark broader than pupil, originating above centre of eye, its lower border extending anterior to eye; a pair of dark-edged white spots anteriorly on snout near upper lip; one more small marks on maxilla; anterior nostril nearly enclosed in a white spot.... 5
5 Perciformes: Percoidei: Cirrhitidae a. Light brown with a broad pale stripe, broadly bordered in black, posteriorly on upper side of body; upper margin of preopercle smooth or with only a few small serrations. Paracirrhites nisus (Tuamotu Archipelago and Pitcairn Group) 5b. Dark brown except caudal peduncle, caudal fin, and extreme ventral part of body posterior to insertion of pelvic fins which are pale (orange in life); no pale stripe on body; upper margin of preopercle finely serrate Paracirrhites bicolor (Caroline Atoll and Tuamotu Archipelago) List of the species occurring in the area The symbol is given when species accounts are included. Amblycirrhitus bimacula (Jenkins, 1903) Amblycirrhitus oxyrhinchos (Bleeker, 1858) Amblycirrhitus unimaculata (Kamohara, 1957) Cirrhitichthys aprinus (Cuvier, 1829) Cirrhitichthys falco Randall, 1963 Cirrhitichthys oxycephalus (Bleeker, 1855) Cirrhitops hubbardi (Schultz, 1943) Cirrhitus albopunctatus Schultz, 1950 Cirrhitus pinnulatus (Bloch and Schneider, 1801) Cyprinocirrhites polyactis (Bleeker, 1875) Isocirrhitus sexfasciatus (Schultz, 1960) Neocirrhites armatus Castelnau, 1873 Oxycirrhites typus Bleeker, 1875 Paracirrhites arcatus (Cuvier, 1829) 1/ Paracirrhites bicolor Randall, 1963 Paracirrhites forsteri (Bloch and Schneider, 1801) Paracirrhites hemistictus (Günther, 1874) Paracirrhites nisus Randall, 1963 Paracirrhites xanthus Randall, 1963 References Randall, J.E Review of the hawkfishes (family Cirrhitidae). Proc. U.S. Natl. Mus., 114: Randall, J.E The hawkfish Cirrhitichthys serratus Randall, a synonym of C. falco Randall. Micronesica, 30(1): / Paracirrhites amblycephalus (Bleeker, 1857), known from a single specimen, 78 mm standard length, from the Sangi Islands (Indonesia) is here treated as a probable synonym of P. arcatus.
6 3326 Bony Fishes Cirrhites pinnulatus (Bloch and Schneider, 1801) Frequent synoynms / misidentifications: Cirrhitus alternatus Gill, 1862; C. spilotoceps Schultz, 1950 / None. FAO names: En - Stocky hawkfish; Fr - Epervier de corail; Sp - Solosolo robusto. Diagnostic characters: Body depth 2.7 to 3 times in standard length. Snout short and blunt. Supraorbital ridge low. Mouth large, the maxilla nearly reaching a vertical at posterior edge of eye; upper margin of preopercle finely serrate. A continuous dorsal fin of X spines and 11 soft rays, deeply notched between spinous and soft portions; a tuft of cirri from membrane near tip of each dorsal-fin spine; anal fin with III spines and 6 soft rays; caudal fin slightly rounded; pectoral fins not reaching a vertical at tips of pelvic fins; pectoral-fin rays 14, the lower 7 unbranched and thickened. Lateral-line scales 39 to 44; 4 rows of large scales above lateral line in middle of body; scales on cheek much smaller than scales on body. Colour: ground colour white but overlaid with squarish dark brown blotches and numerous dark orangish to reddish brown spots, leaving as white 3 rows of 5 or 6 spots of about size of eye on body; ventral thorax and abdomen white; head with reddish brown spots, sometimes conjoined to form irregular bands; median fins with reddish brown spots. Size: Maximum total length 30 cm, commonly to 23 cm. Habitat, biology, and fisheries: This robust hawkfish, the largest of the family in the Indo-Pacific, lives on rocky shores and reef fronts exposed to wave action. Nocturnal; feeds mainly on crabs. Generally caught from shore, usually by hook-and-line. Marketed mostly fresh. Distribution: Occurs throughout the Indo-Pacific region from the Red Sea south to Port Alfred (South Africa) and east to French Polynesia (including Rapa) and the Pitcairn Group; in the western Pacific from southern Japan to the Great Barrier Reef. Remarks: This species is slightly different in colour in Hawaii and appears to have a different count of gill rakers on the upper limb of the first gill arch (5 or 6 in Hawaii compared to 6 or 7 elsewhere). The name Cirrhites alternatus Gill is available in the event a subspecific name is desired for the Hawaiian population.
7 Perciformes: Percoidei: Cirrhitidae 3327 Paracirrhites forsteri (Bloch and Schneider, 1801) Frequent synonyms / misidentifications: Paracirrhites typee Randall, 1963 / None. FAO names: En - Blackside hawkfish; Fr - Epervier à bande noire; Sp - Solosolo de banda negra. Diagnostic characters: Body depth 2.6 to 2.8 times in standard length. Snout short. Mouth large, the maxilla reaching posterior to a vertical at centre of eye; upper margin of preopercle finely serrate. A continuous dorsal fin of X spines and 11 soft rays, notched between spinous and soft portions; a single cirrus from membrane near tip of each dorsal-fin spine; anal fin with III spines and 6 soft rays; pectoral-fin rays 14, the lower 7 unbranched and thickened; caudal fin slightly rounded. Lateral-line scales 45 to 49; 5 rows of large scales between lateral line and spinous portion of dorsal fin; 5 or 6 rows of large scales on cheek, in addition to small scales. Colour: yellowish with a faint longitudinal dark banding following scale rows; usually a broad blackish band (sometimes broken into conjoined spots) on upper side from below middle of dorsal fin nearly to middle of caudal fin; region below black band usually yellow (by virtue of reduced longitudinal banding on scale rows); head, nape, and thorax with small brownish red spots; fins varying from yellowish to pink. Size: Maximum total length 22 cm, commonly to 18 cm. Habitat, biology, and fisheries: A common coral-reef species, often observed sitting on live coral. Feeds mainly on small fishes. Occasionally caught throughout its range. Taken mainly by hook-and-line. Marketed mostly fresh. Distribution: Indo-Pacific, but not known from the Persian Gulf or Gulf of Oman. Tropical and subtropical islands of the Indian Ocean and coast of Africa from the Red Sea to Natal. In the Pacific from southern Japan to New South Wales, east to the islands of Oceania, including Hawaii and the Pitcairn Group.
8 3328 Bony Fishes Paracirrhites hemistictus (Günther, 1874) Frequent synonyms / misidentifications: Paracirrhites polystictus (Günther, 1874) / None. FAO names: En - Halfspotted hawkfish. first phase second phase Diagnostic characters: Body elongate for the genus, its depth 2.8 to 3.2 in standard length. Mouth large, maxilla reaching to or posterior to a vertical at centre of eye. Upper two-fifths of margin of preopercle finely serrate, a slight concavity in margin below serrate portion. A continuous dorsal fin of X spines and 11 soft rays, notched between spinous and soft portions; tenth dorsal-fin spine distinctly longer than ninth; a single cirrus from membrane near tip of each dorsal-fin spine; anal fin with III spines and 6 soft rays; pectoral-fin rays 14, the lower 7 unbranched and thickened; caudal fin slightly rounded, the corners sharply angular; lateral-line scales 48 to 51; 5 rows of large scales between lateral line and spinous portion of dorsal fin; snout almost entirely scaly; 5 oblique rows of large scales on cheek, in addition to small scales. Colour: 2 colour phases, not related to sex, the first greenish on back, densely spotted with black, pale yellow below, with series of dark yellow spots forming stripes following scale rows; an irregular white stripe slightly below midside of body with a few black spots below and occasionally within stripe; head greenish to pinkish grey; fins yellowish. Second (Paracirrhites polystictus) phase grey with numerous close-set dark brown spots on body and a white or pale pink spot slightly smaller than eye on upper side in middle of body; head reddish grey without markings. Size: Maximum total length 29 cm, commonly to 20 cm. Habitat, biology, and fisheries: A shallow-water, coral-reef species generally found at oceanic Islands. Rests motionless on the substratum, often on live coral. Food habits not investigated; probably feeds opportunistically on crustaceans and small fishes that venture near. Generally caught by hook-and-line. Of limited commercial importance. Distribution: Occurs on all the islands of Oceania except Hawaii and Easter Island; also found on the Great Barrier Reef and Christmas Island and Cocos-Keeling Islands in the eastern Indian Ocean. Curiously, it remains unreported from any continental shore or any of the large islands of the Indo-Malayan region.
9 Perciformes: Percoidei: Cheilodactylidae 3329 CHEILODACTYLIDAE Morwongs by J.E. Randall Diagnostic characters: Body oblong and moderately compressed (size to about 45 cm). A single flat spine on opercle; margin of preopercle smooth. Mouth small, terminal to slightly inferior, the adults with thick lips; maxilla largely exposed on cheek; no supramaxilla. Teeth small, villiform, in bands anteriorly in jaws, none on vomer or palatines. Branchiostegal rays 6. A continuous dorsal fin with XIV to XXII spines and 19 to 39 soft rays; anal fin with III spines and 7 to 19 soft rays; caudal fin forked; lower 4 to 7 pectoral-fin rays unbranched, thickened, and prolonged, as much as the distal 50% free of membrane; pelvic fins with I spine and 5 soft rays, their base distinctly posterior to base of pectoral fins. Scales cycloid and small, 45 to 85 in lateral line; dorsal and anal fins with a basal scaly sheath. Vertebrae Colour: most of the species have vertical to oblique dark bands. Habitat, biology, and fisheries: Morwongs are bottom-dwelling fishes, mainly found inshore, though a single species is taken commercially in southern Australia by trawling to depths of 400 m. They feed on a wide variety of small invertebrates which are sorted from the mouthfuls of sand and detritus that they ingest from the substratum. The postlarvae are very deep bodied and extremely compressed (sometimes referred to as the paper fish-stage, and may attain large size. The late postlarval stage of Goniistius vittatus may reach 6 cm total length. A specimen of this species from the northeastern Hawaiian Islands collected from the nest of a white tern was mistakenly described as a new genus and new family, Gregoryinidae, by Fowler and Ball (1924). Most of the species are valuable food fishes, but only a few occur marginally in the Western Central Pacific. Distribution: The Cheilodactylidae is primarily a temperate southern hemisphere family. Australia has the largest number of species, 14; New Zealand has 6 (all shared with Australia), South Africa 5, Chile and Peru 3, and Argentina 1. In the northern hemisphere, there are 3 species in Japan and China and 1 in Hawaii. The group is clearly antitropical in its distribution. Remarks: Randall (1983) classified 8 Indo-Pacific species of Cheilodactylus in the subgenus Goniistius. In a current study of the Cheilodactylidae, Chris Burridge (pers. comm.) has concluded that Goniistius should be elevated to a genus for these species except for Cheilodactylus nigripes that may warrant the description of a new genus. He has reclassified C. ephippiumand C. fuscusin the genus Morwong Whitley. He wrote, I believe that Cheilodactylus should be restricted to 2 South African species, C. fasciatus and C. pixi.
10 3330 Bony Fishes Similar families occurring in the area Of the 4 families of fishes in the superfamily Cirrhitoidea, only the hawkfish family (Cirrhitidae) occurs in the area with the Cheilodactylidae. The species of this family are easily separated by having X dorsal spines compared to XIV to XXII for the cheilodactylids. Key to the species of Cheilodactylidae occurring in the area 1a. Anal-fin rays 16 or 17; dorsal-fin rays 27 or 28; pectoral-fin rays 14 or 15 (usually 15); no bony protuberances anteriorly on head Nemadactylus douglasii (southern Queensland to Tasmania and New Zealand) 1b. Anal-fin rays 8 or 9; dorsal-fin rays 29 to 35; pectoral-fin rays 13 or 14; adults with 2 pairs of knob-like bony protuberances, 1 anterior to eyes and 1 just above upper lip a. Third and fourth dorsal-fin spines subequal; body grey-brown dorsally and on most of caudal peduncle, shading to whitish below, with 2 narrow white bars on caudal peduncle and 2 short oblique white bands below soft portion of dorsal fin Morwong fuscus (southern Queensland to Victoria, east to Lord Howe Island and New Zealand) 2b. Fourth dorsal-fin spine much longer than third spine, the third 3.75 to 5 times in fourth; body whitish with 2 oblique black bands; head whitish, with 2 oblique black bands which converge and join at pectoral-fin base a. Dorsal-fin rays 32 to 35; body depth 2.6 to 2.85 times in standard length; no oblique black band from below eye to chest; pelvic fins not dark brown to black Goniistius vestitus (southern Queensland to central New South Wales, east to Lord Howe Island, Norfolk Island, and New Caledonia) 3b. Dorsal-fin rays 29 to 33; body depth 2.5 to 2.6 times in standard length; an oblique black band from below eye to ventrally on chest; pelvic fins dark brown to black.... Goniistius vittatus (Hawaii, Lord Howe Island, Kermadec Islands, and New Caledonia) List of species occurring in the area Goniistius vestitus (Castelnau, 1878) Goniistius vittatus (Garrett, 1864) Morwong fuscus (Castelnau, 1879) Nemadactylus douglasii (Hector, 1875) Cirrhitidae References Gomon, M.J., G.C.M. Glover, and R.H. Kuiter (eds) The fishes of Australia s south coast. Adelaide, State Print, 992 p. Randall, J.E A review of the fishes of the subgenus Goniistius, genus Cheilodactylus, with descriptions of a new species form Easter Island and Rapa. Occ. Pap. B.P. Bishop Mus., 25(7):1-24.
11 Perciformes: Percoidei: Cepolidae 3331 CEPOLIDAE Bandfishes by W.F. Smith-Vaniz Diagnostic characters: Small (to about 40 cm), moderately to noticeably elongate fishes with compressed, tapering body and lanceolate caudal fin. Head short, with blunt snout. Eyes relatively large and high on head. Mouth large, oblique; upper jaw broad at end, without supramaxilla, and extending to below posterior margin of eye; a single row of slender, slightly curved teeth in jaws with an inner cluster of teeth at symphysis in some species. Dorsal fin continuous, with 0 to IV flexible spines and 21 to 89 segmented rays; anal fin with 0 or I spine and 13 to 102 segmented rays; caudal fin lanceolate, middle 9 to 15 rays branched; pelvic fins positioned below or slightly anterior to pectoral fins, with I spine and 5 segmented rays; outermost segmented pelvic-fin ray unbranched or weakly branched, 4 inner rays distinctly branched. Lateral line high on body, close to dorsal-fin base, terminating posteriorly near end of fin; lateral-line tubes or canals on scales not embedded in skin. Scales cycloid (smooth) or with crenulate margins, relatively large to minute. Colour: in life red or pink; most species have a distinctive dark stripe on the membrane (usually hidden) connecting the premaxillary and maxillary bones of the upper jaw. Sphenanthias macrophthalmus Owstonia maccullochi Sphenanthias sibogae Acanthocepola limbata Habitat, biology, and fisheries: Relatively uncommon fishes taken by trawls in shallow to deep depths (to at least 475 m). Bandfishes occur on level bottom, sand or mud substrates where they live in burrows, which they excavate themselves. Not of significant commercial importance, but consumed by the local population in some areas. Similar families occurring in the area The combination of a lanceolate tail, large oblique mouth, and the arrangement of the pelvic-fin rays, consisting of I spine and 5 segmented rays (the outermost ray unbranched or weakly branched and the inner 4 branched), will distinguish the bandfishes from all other families. Additional distinguishing characters of the superficially similar Opistognathidae are the following: Opistognathidae: dorsal-fin spines X or XI (0 to IV in Cepolidae); caudal fin rounded (moderately to strongly lanceolate in Cepolidae); pelvic fins with 5 segmented rays, the outer 2 unbranched. Opisthognathidae
12 3332 Bony Fishes Partial key to the species of Cepolidae occurring in the area Note: the taxonomy of this family is very poorly known and even generic definitions and limits are not well established. Following traditional usage, Sphenanthias Weber, 1913 is tentatively recognized as distinct from Owstonia Tanaka, 1908, solely on the basis of a different lateral line configuration (see key). However, some species of Sphenanthias appear to be more closely related to species of Owstonia than to S. sibogae, the type species. A number of undescribed species belonging to the subfamily Owstoniinae occur in the area and at this time it is not possible to provide a reliable key for their identification. 1a. Total dorsal-fin elements (i.e. spines plus soft rays) 71 to 89; last soft ray of dorsal and anal fins narrowly to broadly connected to caudal fin by a membrane..(subfamily Cepolinae) 2 1b. Total dorsal-fin elements 23 to 27; last soft ray of dorsal and anal fins not connected to caudal fin by a membrane....(subfamily Owstoniinae) 6 2a. Preopercular margin with spines; cheeks scaly b. Preopercular margin without spines; cheeks naked a. Middle 8 to 10 caudal-fin rays branched (segmented caudal-fin rays 7 in upper lobe, 6 in lower lobe); vertebrae Acanthocepola abbreviata 3b. No branched caudal-fin rays; vertebrae a. Segmented caudal-fin rays 6 or 7 in upper lobe, 6 or 7 in lower lobe (total 12 to 14); vertebrae Acanthocepola krustensternii 4b. Segmented caudal-fin rays 5 in upper lobe, 5 in lower lobe (total 10); vertebrae Acanthocepola limbata 5a. Dorsal fin with III spines and 54 to 57 soft rays; anal fin with I spine and 50 to 53 soft rays; vertebrae Cepola australis 5b. Dorsal fin with III spines and 65 soft rays; anal fin with 62 soft rays, anal-fin spine absent; vertebrae Cepola schlegelii 6a. Lateral line continues anteriorly, forming loop in front of dorsal fin Owstonia spp. 6b. Lateral line does not form loop in front of dorsal fin Sphenanthias spp. List of species occurring in the area Acanthocepola abbreviata (Valenciennes in Cuvier and Valenciennes, 1835) Acanthocepola krustensternii (Temminck and Schlegel, 1845) Acanthocepola limbata (Valenciennes in Cuvier and Valenciennes, 1835) (Cepola indica Day, 1888 and Acanthocepola cuneatus Smith, 1935 are both considered to be synonyms) Cepola australis Ogilby, 1899 Cepola schlegelii Bleeker, 1854 Owstonia dorypterus (Fowler, 1934) Owstonia maccullochi Whitley, 1934 (may be a junior synonym of O. totomiensis) Owstonia totomiensis Tanaka, 1908 Sphenanthias grammodon (Fowler, 1934) Sphenanthias macrophthalmus Fourmanoir, 1985 Sphenanthias nigromarginatus Fourmanoir, 1985 Sphenanthias sibogae Weber, 1913 (S. pectinifer Myers, 1939 appears to be a synonym) References Mok, H.-K Osteological evidence for the monophyly of Cepolidae and Owstoniidae. Japan. J. Ichthyol., 34 (4): Smith, J.L.B New and interesting fishes from deepish water off Durban, Natal and southern Mozambique. S. Afr. Ass. Mar. Biol. Res. Invest. Rep., 19:1-30. click for next page
click for previous page Groupers of the World 281 Literature: Walford (1937). Remarks: Rosenblatt and Zahuranec (1967) question the place of origin of the holotype: Although the type locality of M. xenarcha
SCTB17 Working Paper FTWG INF 4 A HANDBOOK FOR THE IDENTIFICATION OF YELLOWFIN AND BIGEYE TUNAS IN BRINE FROZEN CONDITION David G. Itano Pelagic Fisheries Research Program JIMAR, University of Hawaii USA
click for previous page 1740 Bony Fishes Suborder ZOARCOIDEI ZOARCIDAE Eelpouts by M.E. Anderson, South African Institute for Aquatic Biodiversity, South Africa Diagnostic characters: Small to medium-sized
click for previous page 1898 Bony Fishes PARALICHTHYIDAE Sand flounders by T.A. Munroe, National Marine Fisheries Service, National Museum of Natural History, Washington D.C., USA Diagnostic characters:
Copyright Australian Museum, 2004 Records of the Australian Museum (2004) Vol. 56: 147 158. ISSN 0067-1975 Five New Fish Species of the Genus Alabes (Gobiesocidae: Cheilobranchinae) BARRY HUTCHINS* AND
Bull. Kanagawa prefect. Mus. (Nat. Sci.), no.38, pp. 107-112, Mar. 2009 107 Two New Species of the Genus Trimma (Perciformes: Gobiidae) from Japan and Palau Toshiyuki SUZUKI 1) & Hiroshi SENOU 2) Abstract.
Vol. 45, S1: 683-693, diciembre 2010 Article Review of the southern hemisphere fish family Chironemidae (Perciformes: Cirrhitoidei) Revisión de los peces de la familia Chironemidae (Perciformes: Cirrhitoidei)
OCCASIONAL PAPERS OF THE MUSEUM OF ZOOLOGY UNIVERSITY OF MICHIGAN THE FIRST ICNOWN BLIND FISH OF THE FAMILY CHARACIDAE: A NEW GENUS FROM MEXICO AMONG the interesting fish novelties being introduced from
click for previous page 2004 Bony Fishes Order LOPHIIFORMES LOPHIIDAE Anglerfishes (monkfishes) by J.H. Caruso Diagnostic characters: Head and anterior part of body usually depressed and very broad, posterior
chapter 4 GROUPERS 4 GROUPERS Introduction and species identification Groupers (class Actinopterygii, order Perciformes, family Serranidae, subfamily Epinephelinae) are classified in 14 genera of the subfamily
SPECIES DESCRIPTION Picture German Cockroach Cockroach egg case & nymphs 12-15mm (Adult Stage)Mid to dark brown body. Lighter area with 2 dark bars behind head. Wings. Very long antennae. Young are smaller
click for previous page 44 Geographical Distribution : Eastern Pacific from Baja California, Mexico (27ºN) to Valparaiso, Chile (ca. 32ºS) (Fig. 77). As Manning (1970:868) pointed out, the records from
A new species of deep-sea catshark (Scyliorhinidae: Bythaelurus) from the southwestern Indian Ocean DAVID A. EBERT Pacific Shark Research Center, Moss Landing Marine Laboratories, Moss Landing, CA 95039,
West Virginia Trees Basic Tree Identification For FFA Forestry Contest Developed in cooperation with the West Virginia Division of Forestry and the Tyler County FFA Chapter. Identification of trees base
Genus Vol. 13 (3): 409-415 Wroc³aw, 30 IX 2002 A new species of Myrmarachne from Kenya (Araneae: Salticidae) WANDA WESO OWSKA 1 and KATHRYN SALM 2 1 Zoological Institute, Wroc³aw University, Sienkiewicza
information leaflet Insect identification sheet To enable you to identify the insect you have in your home, we have provided some identification sheets. These depict the most common pest species that you
AUSTRALIAN MUSEUM SCIENTIFIC PUBLICATIONS Bayly, I. A. E., 1971. A new species of Kelleria (Copepoda: Cyclopoida) from brackish water in Victoria. Records of the Australian Museum 28(6): 111 116. [27 September
Fishy Adaptations Adapted from: Fashion a Fish in Project Wild Aquatic Education Activity Guide. The Council for Environmental Education, 1992 Physical Structure Grade Level: Basic Duration: 45 minutes
2012-2013 ORAL BIOLOGY DEVELOPMENT AND GROWTH OF THE MANDIBLE Ass. Prof. Dr. Heba M. Elsabaa Development and Growth of the Mandible DEVELOPMENT OF THE MANDIBLE The Mandible Is the largest and strongest
Available online at www.scholarsresearchlibrary.com Annals of Biological Research, 2015, 6 (2):10-14 (http://scholarsresearchlibrary.com/archive.html) ISSN 0976-1233 CODEN (USA): ABRNBW Morphometric and
click for previous page Sharks of the World, Vol. 2 161 Orectolobus sp. A Last and Stevens, 1994 Fig. 125 Orectolobus sp. A Last and Stevens, 1994, Sharks Rays Australia: 128, pl. 26. Synonyms: None. Other
Exploring the World Classroom Activity The Classroom Activity introduces students to the context of a performance task, so they are not disadvantaged in demonstrating the skills the task intends to assess.
click for previous page Pleuronectiformes: Achiridae 1925 ACHIRIDAE American soles by T.A. Munroe, National Marine Fisheries Service, National Museum of Natural History, Washington, D.C., USA Diagnostic
click for previous page 670 Bony Fishes Order ACIPENSERIFORMES ACIPENSERIDAE Sturgeons by W.B. Scott, Kingston, Ontario, Canada Diagnostic characters: Body heavy, elongate and subcylindrical in section.
Abstract Review of the Indo-Pacific labrid fish genus Hemigymnus JOHN E. RANDALL Bishop Museum, 1525 Bernice St., Honolulu, HI 96817-2704 USA. E-mail: firstname.lastname@example.org The labrid fish genus Hemigymnus
Advanced MakeUp Tips by Kathy Whittington Applying Eye Shadow Eyes are the windows to your soul. The right make up can add to the beauty of your eyes if you know how to accent the positive and minimize
Alebion echinatus Capart from Japanese Waters, with Observations on the Newly Found Male Form 1 SUEO M. SHIIN0 2 THE HAMMERHEAD SHARK, Sphyrna zygaena (Linne), was reported by Wilson (1932) as infected
Missouri Snakes information from Mo. Dept. Of Conservation Environmental Science 2010-2011 Black Rat Snake Elaphe obsoleta Black Rat Snake Elaphe obsoleta Generally shiny black, but some individuals show
Educational Material Bony Fish Anatomy Worksheet Teacher Information This activity is designed to be team-taught by the classroom and art teacher. Use this guide in conjunction with fish-related art to
Key to the Snakes of South Australia Austrelaps labialis Mark Hutchinson & Ian Williams Version: 24 October, 2014 KEY TO THE SNAKES OF SOUTH AUSTRALIA ELAPIDAE, PYTHONIDAE 1. a. Eyes rudimentary, visible
Lab 1: External Anatomy & Taxonomy Goals: 1. Learn the basic external anatomy of fishes. 2. Learn how to identify fish species. Tasks: 1. Choose a fish. 2. Identify all major external structures of your
A Patient s Guide to Lumbar Spine Anatomy 228 West Main St., Suite D Missoula, MT 59802-4345 Phone: 406-721-3072 Fax: 406-721-2619 email@example.com DISCLAIMER: The information in this booklet is compiled
1944] Three Species o[ Coleosoma ]rom Florida 51 THREE SPECIES OF COLEOSOMA FROM FLORIDA (ARANE/E; THERIDIID.E) x BY ELIZABETH B. BRYANT Among material recently sent the Museum of Comparative Zoology by
Zootaxa : 1 8 (2004) www.mapress.com/zootaxa/ Copyright 2004 Magnolia Press ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) Description of two new species of the catfish genus Trichomycterus
click for previous page CHIMAERAS by D.A. Didier, Academy of Natural Sciences, Philadelphia, Pennsylvania, USA 592 Chimaeras TECHNICAL TERMS AND MEASUREMENTS total length spine frontal tenaculum lateral-line
Platnick, N. I., and M. U. Shadab. 1977. A new genus of the spider subfamily Gnaphosinae from th e Virgin Islands (Araneae, Gnaphosidae). J. Arachnol. 3 :191-194. A NEW GENUS OF THE SPIDER SUBFAMILY GNAPHOSINA
Activities in Museums and Aquaria 143 A Trip to the Museum Level 1-5 Key questions What do marine organisms look like? How are they different from terrestrial animals? How do they adapt to their environment?
GreenCustoms Knowledge Series No. 11 Identification of snake skins Developed by the CITES Secretariat Questions to answer What is snake skin used for? How to identify snake skin? What are the common CITESlisted
Source: Klemm, E.B., S.A. Reed, F.M. Pottenger, C. Porter, T.W. Speitel. 1995. HMSS The Living Ocean. Honolulu, HI: Curriculum Research & Development Group, University of Hawaii. Pp. 14-21. Adapted and
PowerPoint Lecture Slides prepared by Barbara Heard, Atlantic Cape Community College Ninth Edition Human Anatomy & Physiology C H A P T E R 7 The Skeleton: Part B Annie Leibovitz/Contact Press Images Vertebral
OCCASIONAL PAPERS OF THE MUSEUM OF ZOOLOGY UNIVERSITY OF MICHIGAN ANN ARBOR, MICHIGAN UNIVERSITY OF MICHIGAN PRESS NOTES ON THE GENUS PLECTROHYLA, WITH DESCRIPTIONS OF NEW SPECIES TIIE generic and specific
United States Department of Agriculture Agricultural Marketing Service Fruit and Vegetable Division United States Standards for Grades of Lettuce Fresh Products Branch Effective December 1, 1975 (Reprinted
Snakes of Massachusetts Of the roughly 3,000 known species of snake found worldwide, only 15% are considered dangerous to humans. Luckily here in Massachusetts we only have two types of dangerous venomous
CHAPTER 10 RESTS AND DEFINITIONS A REST is any rigid part of an RPD framework which contacts a properly prepared surface of a tooth. A REST PREPARATION or REST SEAT is any portion of a tooth or restoration
BULLETIN OF THE U N IV E R SIT Y OF UTAH Vol. 35 March 15, 1945 No. 17 New Mites in the Family Caeculidae B Y STANLEY M ULAIK BIOLOGICAL SERIES, Vol. VIII, No. 6 PUBLISHED BY THE UNIVERSITY OF UTAH SALT
FIELD RECOGNITION OF THE LARVAE OF NATIVE COCCINELLIDAE, COMMON TO THE POTATO FIELDS OF AROOSTOOK COUNTY R. H. Storch Department of Entomolgy, University of Maine TECHNICAL BULLETIN 43 MAINE AGRICULTURAL
THE RAFFLES BULLETIN OF ZOOLOGY 2004 THE RAFFLES BULLETIN OF ZOOLOGY 2004 52(2): 455-466 National University of Singapore TWO NEW FRESHWATER SPECIES OF THE GENUS JESOGAMMARUS (CRUSTACEA: AMPHIPODA: ANISOGAMMARIDAE)
Week 1 1. What US city has the largest population? 2. Where is Aachen? 3. What is the capitol of Florida? 4. What is the longest mountain range in Spain? 5. What countries border Equador? Week 2 1. What
17.12.2015/EN FEDERATION CYNOLOGIQUE INTERNATIONALE (AISBL) SECRETARIAT GENERAL: 13, Place Albert 1 er B 6530 Thuin (Belgique) (VALID FROM 01/01/2016) FCI-Standard N 200 PICCOLO LEVRIERO ITALIANO (Italian
A new araneid genus from the Galapagos Islands (Araneae: Araneidae) Herbert W. Levi ABSTRACT Contrib. Nat. Hist. 12: 893 898. A new araneid, Galaporella thaleri sp. nov., similar to species of the genus
CLASS: CIRRIPEDIA BARNACLES They are highly modified crustaceans. Along the shore they live permanently attached to rocks in the balanoid zone. Can also attach themselves to other solid substrata, e.g.
12.01.2011/EN FEDERATION CYNOLOGIQUE INTERNATIONALE (AISBL) SECRETARIAT GENERAL: 13, Place Albert 1 er B 6530 Thuin (Belgique) FCI-Standard N 122 LABRADOR RETRIEVER M.Davidson, illustr. NKU Picture Library
A new fish, Peristedion nesium (Scorpaeniformes: Peristediidae) from Isla del Coco, Costa Rica William A. Bussing Escuela de Biología & Centro de Investigaciones en Ciencias del Mar y Limnología (CIMAR),
Lectures of Human Anatomy Vertebral Column-I By DR. ABDEL-MONEM AWAD HEGAZY M.B. with honor 1983, Dipl."Gynecology and Obstetrics "1989, Master "Anatomy and Embryology" 1994, M.D. "Anatomy and Embryology"
Name: Maui Ocean Center Learning Worksheets Fifth Grade Our mission is to foster understanding, wonder and respect Hawai i s for marine life. Based on benchmarks SC.5.5.1, SC. 5.3.1 1 Create a Food Chain
Lab 5 Overview of the Skeleton: Classification and Structure of Bones and Cartilages Exercise 9 The Axial Skeleton Exercise 10 Overview of the Skeleton Locate the important cartilages in the human skeleton
The Blue Velvet Angelfish Centropyge deborae sp. nov., a New Pomacanthid from the Fiji Islands, Based on Genetic and Morphological Analyses Kang-Ning Shen 1, Hsuan-Ching Ho 2,3, and Chih-Wei Chang 2,3,
Communities, Biomes, and Ecosystems Section 1: Community Ecology Section 2: Terrestrial Biomes Section 3: Aquatic Ecosystems Click on a lesson name to select. 3.1 Community Ecology Communities A biological
United States Department of Agriculture Agricultural Marketing Service Fruit and Vegetable Programs Fresh Products Branch United States Standards for Grades of Grapefruit (Texas and States other than Florida,
An Identification Guide for North Texas Snakes Text & Photos - Michael Smith There are many species of snakes living in north Texas, and most of them are harmless. A few species are, and even those tend
UNIT 3 SALMON ANATOMY SALMON ANatOMY Overview The class identifies the body parts of a fish and compares them to human body parts. They see how fish shape, skin, scales and gills help salmon live in water.
Simplified Positioning for Dental Radiology Prepared by: Animal Dental Care Tony M. Woodward DVM, Dipl. AVDC 5520 N. Nevada Ave. Suite 150 Colorado Springs, CO 80918 (719) 536-9949 firstname.lastname@example.org www.wellpets.com
Article 56.9,33N XLI.- PRELIMINARY DIAGNOSIS OF AN APPARENTLY NEW FAMILY OF INSECTIVORES. BY H. E. ANTHONY. PLATE XXIII. In July, 1916, while searching for remains of fossil mammals in Porto Rico, in accordance
Araştırma Makalesi/Research Article A Morphological Study On Endemic Malabaila lasiocarpa Boiss. (Apiaceae) From Bingol (Turkey) Ömer KILIÇ 1 Abstract- In this study morphological characters of Malabaila
50 FAO Species Catalogue for Fishery Purposes No. 4, Vol. 1 2. CHAMBERED NAUTILUSES by Patrizia Jereb Living nautiluses are limited to a few species belonging to 1 family and 2 genera: the sole survivors
Classification of Malocclusion What s going on here? How would you describe this? Dr. Robert Gallois REFERENCE: Where Do We Begin? ESSENTIALS FOR ORTHODONTIC PRACTICE By Riolo and Avery Chapter 6 pages
Bandwinged Grasshoppers Subfamily Oedipodinae The bandwinged grasshoppers are usually heavy bodied and bear enlarged hind legs. The head of these grasshoppers often appears enlarged and broadly rounded.
Cranoglanis henrici (Vaillant, 1893), a valid species of cranoglanidid catfish from Indochina (Teleostei, Cranoglanididae) Heok Hee NG Fish Division, Museum of Zoology, University of Michigan, 1109 Geddes
A Beginner s Guide to Butterfly Identification Common Name Color(s) Key Feature(s) from one wing tip to the other Zebra Swallowtail (SWT) White with stripes Very long tails zebra striped 2½ -4 Moist woods
Female Dog Urinary Catheterization 1 of 6 Placement of an indwelling urinary catheter in female dogs Bernie Hansen DVM MS North Carolina State University College of Veterinary Medicine Materials Needed
Excerpt from Verlag Dr. Friedrich Pfeil ISSN 0936-9902 Ichthyological Exploration of Freshwaters An international journal for field-orientated ichthyology Volume 25 Number 1 This article may be used for
CHEM 107 Hair handout. 3-22-05 and 3-24-05 Basic Structure of Hair A hair can be defined as a slender, thread-like outgrowth from a follicle in the skin of mammals. Composed mainly of keratin, it has three