INDTAN AOltTOULTURAL. RESEARCH INS'l'ITUl'E, NEW DELHI. I. A. R ]\igipg-sl-6 Al~/oL ,OOO.

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1 INDTAN AOltTOULTURAL RESEARCH INS'l'ITUl'E, NEW DELHI. I. A. R ]\igipg-sl-6 Al~/oL ,OOO.

2 LIST OF SUGAR-CANE'l~SECTS A synonymic Catalogue of the Sugar-cane Insects and l\1ites (If the Wadel, and of their Insect Parasites and P"ediltl'Jrs, arranged systematically. By HAROLD E. Box, F.R.E.S., F.R.G.S. LONDON: COMMONWEALTH INSTITUTE OF ENTOMOLOgy 41, ~QlJEEN'S GATE, LONDON, S:W.7 " 1953 All Rights Reserved Imllilllij;I~II~IIIIIIIIIIII 1111,liA.RI lsmotla1'l Agrl~!rearcrr.rnsun:ltie New!).e,w. (I,bk)

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4 ipl'ifge UST OF SUGAR-CANE INSECTS By :i!l<i!jl. Bale, F,R.ll!.R" F':a.a,s., ( I ERRATA 6 (Aildotll$10?'n'H)8((J1t~UJ)' for HatllnychiUIJ road PlatynyehUfI 10 (De1'17l,OZepida I),lbohirt~bm) - for Trisiloa read T~8&/'loa 20 (Amplipalpa (see OediOpalpIL) - transfer to Hispirlae, bolow 23 (TeLtoo~a) - tor kreffti read (I!!LgitHl k'rejfli, Macl-, 28 (Chilo)" 01' (/emindtal~8 mall (/ernm~nota.l(8 31 (Gi1'dhClJ!"ttJ, ta1,1'omma) - for (Jlnt()torrente/lu~8 J:'E'ad Chi-Ia torrentellllll C01ltiodes a('cepta) - delete I, (2)" after J-Iorogcucs 44 (AeneoZr:umia (Tomall'pis) - itlsert "varut, F. continued" in liue llelow 61 (IIwl'ogllJphns) - fol' C'I'iZL'/)O}'U8 remi orizlv()ru.$ 67 (Hypel'a8pia je8tiva).~ for D01l1illCan read Dominican 7B (K1'tt'(}ena hoptazonat;) - for un1-formu;[ read ~i!h.f(ll'ntis 74 (Stau,rolwdoctolnlAl 71UZ,1t'l'iti~) -!ov l;ipsamia vutcna ro l>d,'>'('sa11luf, dalamisits 77 ('1'l'irllogpilu8 pl/pwora) " tor Se8amta 1}utcrta read Sesamlla ('ala1l!lrtfr 78 (ljraehymel'iq; av{lta) - for Pam,ogwtna refl,cl Nyc{eUul< 79 (Telerw'l/tlUi bcnejlclens) - for BeBamuJ, Vlde'ria road Scsamfa calw14sti8 80 (Canvp8onuJris a'l'/l!h~lata) for e'll1jm!8a read e21lan8a 84 (Blondelza armigera,) for jl'1{ltperaa road j'l'u!]'i1)er(l({ 815 (JIypn,~tena dtait'u('ae) - fol' JiM raeae roaa (lri8lja Curl', 87 (ParMlzel'('8W, cla1'iljal;pis) for Me'\.ico t read Mexico IND1~X TO COUNTRIES B.wbac1of, -.for 52, 54 read Brazil ~/M' 5U read 54, 56; ad(! 86 Ohina -' jo}' 6il 1'I'ad 62 Cuba - j01' 11, l.j-ib react Dmnllllca - de/de 76 DCIl!II11ican Rellublw - add 76 EI!ypt - jar 55 read 54, 56 FI)i Islancl" - jo)' 24, 25 r(all Haiti - (til(l Hfi Hit wl1lian I~lll,llllf, - [01' 1i4,,, 59 reo a 54- ')9 Jamaica jar 60-63?'Carl BO, 62, 63; ad(l 66 anrl 69 J apau - lele[r flli, 09 1\lexico - for 59, 61, 1)2, read Nicaragua "/01' 25 read 26 PCI'll - for Jil, read 1il-lti Philippine IslandS,for 66,, ' 77 reatl Queenslancl - add 43 Ryukyu Islanll"!. jo)' J'e(~d 40, 42 Ht, Cl'oix /O?' read 13, 14, J f:i ; 1M' 32 read,50 Trinic1ad - add )0 Vcnezuela -.for 50,., 6;1 l'ead 50.54, 59 6S AgeZl1w - for 48? ead 45 Ar!1'lIplllW - acld 66 Aluctta - add 33 Om'omusia, - fm' 85 read 84 LcpWopte1'a - Jar 24 1'ead 134 SYSTEMATIC INJi1'EX Nymphalit1ae - delete thz8 entrjj Podoth?'IP8 - add 56 Pnon1UJ - for 19 j'ea{l 18 Scoliidae - /01' 80 1'ead XLph1,lto'YI - acl{t 89

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6 "~XVIII. Erllca minima e rtlbro [,{ca. The T;Jlonn eating the Sugar Canes. )i This is Dot over one third of an Inch long, and not fa thick as a t'-lens Qui11, reddiih brown of Colour. It is convey'd in at firft by a round Hole made in the green Sugar Cane, and thence makes a two or three Inches long, rounel, red Cavity the length of the Cane whence when per feet: it comes out. It is in Probability a Sort of a Buttedly, Moth, Beetle or WeevilI, which ihrufl:s in the Egg, tben 'tis hatch'd and feeds on this Cane till it be 'ready to be turn'd to an Aurelia, whence it comes out and leaves a greyilli Skin which I have often found in the Canes fo fpoil'd. The Canes fa eaten are not fit to make Sugar, and therefore are ground to make Rum, or given to the Hogs to feed on. Sometimes Arm eat into the Canes, and ha ve their Young in them. An ondy rcferem:o to SUWLl'-CllllEl Entmnnlogy in JmIlltiefl from Sir Hans Sloane's "A Voyage t,o j,110 I,,;lands ]\Iadom, Rwlxlllos, NioVllS, St. Christophel's I1Ill!,J lima ina ; with tho Natul'fII l-lihtolt of til(> Herbs lind Tl'fmS, Four-footed Beusts, Fishes, Birds,- Insects, IteptiloR, &c, of tho lttrt of t.hose It;lnnds" Vol. II, p_ 220, Loudon, 172;', (Rep1'Oducc(Z by kind pcl'lnjssion of the Royal Entollwloaz'cal Bodety oj London,)

7 This work is dedicated in gratitude to DR. F. X. WILLIAMS Entomologist of the Experimental Station of the Hawaiian Sugar Planters' Association and Author of the incomparable HANDBOOK OF THE INSECTS AND OTHER INVERTEBRATES OF HAWAIIAN SUGAR CANE FIELDS who, just thirty years ago, when staying at Plantation Blairmont, Berbice, British Guiana, inspired a very young entomologist to take a serious interest in the Insects associated with the Sugar-cane. Port of Spain, Trinidad, B.W.I October 1952

8 UONTENrp~ Introduction Acknowledgements...;... 3 Part I-Sugar-cane Insects... 5 Part II-Parasites and Predators....6:3 Bibliography of principal vvol'ks cunsulted... no Index to Countries Systematic Index

9 INTIWDUCTION THIS List has been prepared with the object of bringing together in a convenient form all avaihlble records of sug Ll'-Cane insects, their parasites [lud insect predators, and also that it may serve as a working index to the names used in the litemture of sugar-cane entolilology_ Special attentioll has therefore been given to the varions altcrnative generic and triviallhlmes (not necessarily :>Yllonyms), including en'oneollk names, under which each insect Ims been recordecl. The name ultiumtely selected for a species is intended to conform fls far as possible with the most recent tltxonomic works, aud to further thir end the opinions have frequently been sought of recognised authoritic8, who lmve most obligingly checked fmc! revised the nomenclature ill their l'erpecti\~c grolll's, The greatest. mtl'c has heon taken to a.void final reference to [l speciei': by IllOl'C thrm one llmne, nihl t.he Ol'oss-J.'derencoR shonld olutble any Rpeeies readily to be traceclulldor any of t.ho nn.mer which have been applied t.o it as 11 imgarcalle insdct, flo that furt.her infol'nmtion concerning it may be obtrtined from the appropriatc sonrce,,' in the litcrn.tuj.'e. All attempt has been ma.do 11,1. completeness to the time of IHtllcliug ill till' fina.l t,vpescl'ipt (DC"colllhcr, 105:2). The eompilcr's.first working l.h1nis for the pl'cf;ellt List wnh G, 'V. Kirlmld:r'H BibliorJl'(!]Jhy oj 8/1{/(l1' Cane Ent01lwloY1J, published at HOllolulu ill 1900, a lllllht carefully prepared work from which very few of the eaxly referenee:-;, and certainly none of imporkmce, wore omitted, This was followed by 1\;n exlmm:tive searell through the thirty-nine completcd volumes ( ) ami clll'rent ihsues (Hl52) of the Review oj Applied lentomolo(fl/, 8c1'ies A, and an ex[lmination of tho regional lists mulled in the Bibliography of principnl works consulted. At least ninety per cent. of the reeoitls rue taken froill the a.bovd-lllentioned publica,tiollh, must of the r ldereomingfroltlnumerousshorterandmorerestrictedpapm.h.chiefly those of a systematic llitture trenting of illdividwll groups of il1:;ect.~, A munljei' of additional reuorcis, Inany t,j! them unpublished, have been furnished by the specia,listh and other entomologists who have so.kindly COllllbol'a.terl ill the compilation of thih List. It may be accepted, therefore, that It serious elldea >'OUI' 1111N oeen mn.de to consult if not all then at least the prillcilml SOUl'CeH whence illfol'ilul,tion on the 1mbject might be fol'theoming, A work of this nature, however, is )lot likely to lje.fmc from mistn.kes, nnd it is sij.wcrely JlOpml that olltoluologists lising it will Hot he8itate to bring to Ow notice of the ant.hor any errore: or omissions tlhtt may be found, No a,ttempt has bcoll made to indim1t,e the NtatuH of (LIlY inrect, i.e, whether it be a major pest or a rare speeie:> oceasiollally feeding 011 :mgnr-cltlw, bllt where there is any doubt of the status of an insect, as ill the eatle of some" etlsual " sl'ccieh, then it is included. NOl'ma.lly, however, the criterion foj' inclwlioll in the IH'oRont List is tha.t the insect feeds on the ti8rmes, though not nec:e8sttl'ily the living tishilelh, of thc sugar-cane plant (8acclwru7/1, officl:nrtl'um L., cortain cultiv11tml 81l,ccha1'1.lil/. specie:;;, and hybrids among these), General saproplhlgollh insects, thllt fee(l ill tlhy sort of illjured or decomposillg vegetl1hle matter, are excluded, but seoolhhlry specie,; which fced regularly on Hugnl'-Cane already injured by primary pests, ltre im:ludod. No aceonnt is taken of the many kindr of insects which Imve aclltptod tholllselves to living in " borer " tunnels, even though in 11 stricti :-;el\re they belong to the fi1l111a of the sugar-c11ne plant, [mel neither are inelulled here tho various!tilts which a,t;tend and foster aphids, coccids a,ud other ilolllopterolls insect;; on sugar-cane, For collveniencll the reeol'llecl mites on RugUl'-emw are inehlcled. VariouH local JistS-fl.llrl some o.f wider llpplicf1tioll!tlso-inelucle llllillbers of insects which, though collecteel in Hugal'-Cane fields and sollletimes permanently associated with the habitflt because they feed regularly on mme-field woeds, fwe not (lirectly coneerned with the sugal'-c<1ne plaut in any way, 01' howe so Blight all 1

10 a:-;xocintion with it that, they may be disregarded. Indeed, some mo(lc!'ll checklists of sugar-cane insects have included a considerable nnmber of specim; with no olaim whatever for inclusion other than that specimens were collected perching on the foliage or happened to have been observed in a field of sugar-cane. Nevertheless, due consideration has been given to each sel)arate case before a recorded sugar-cane insect has been finally rejected from the prehent List, and in the majoril;y of cases the rejection has been made after consultation with the local entomologists best able to decide. Part I of the List presents the sugar-cane int';ects of the world and Part II their parasites and insect predators. The sequence of orders and families in both pa,rts follows in the nmill that adopted ill. the latp:-;t. volulllc:'; of the Zoological Record, part Insecta. Genera are arranged in a.1phabctieal order,dthin each family awl the trivial Hallles ill alplmbetical order wit.hill each genus. The ThYHH.lloptel'a, IsopterH and the smaller orders are not classified into families. SVllOllYIl1S, erroneous name8. and names not ill currcnt w;e, are given in braeket".. 0 Particular care has been given to the names of countries; with fr."w exceptions these are in accordance with The O:tford Atlas (Oxford University Press, Hl51), and are arranged in the following geographiclll order: Africa, Asia., AU8tralasia.. Oceania, America, genemlly from west to ea"t in the Old 'World and from north to south in the New World. The Index to Countries provides a ready mealls for the compilation of loca'! lists for indidduaj and neighbonrillg territories. Records llot published to the uate of i88110 of this List (1,1'(1 indica.ted by the sign: t. \Vhere no ot,her person i~ indicated, the present writei' i~ responsible for the record in the count.ry named. While, as stated above, every effort has been matle to make this List Rccnrate and complete, it is hoped to publish witahle supplellelltll from time to time, as ncw records or changes in nomenclature may require.

11 ACKNOWLEDGEMEN'!'S This work would not have been possible without the aid of those who have given generously of their time to collaborate in its prepal'l1tioll., lwd the compiler is under a very deep obligation to the distinguihhed entomologists named below for their unfailing response in checking and revising either systematic lists concel'lling special groups or r,_jgional lists of sugar-cane insects which were sent to them, as well as for other help,. Dr. J. G. Betreru, Holland (Hymenoptera: Scoliidae) lvir. F. A. Bianehi, Honolulu ('l'hysanoptora) Dr. S. C. Bruner, California (Sugar-eane insects of Cuba) Dr. E. A. Chapin, Wlti"lhillgton (Coleoptera: Coceinellidae) Dr. W. E. ChinE" London (Hemiptera: Jassidae).Mr. J. F. Gates Clarke, Washington (Micro-Lepidoptera) lvir. J. P. Doncaster, London (Hemiptera: Aphididae) Dr. F. I. van Emeien, London (Diptera) Brigadier W. H. Evans, C.S.I., C.LE., D.S.O., London (Lepidoptera: Hesperiidae) Mr. R. G. EelllHLh, Trinidad (Hemiptera: Fulgoroidea (Cixiidae to Lophopidae) and Cercopiuae; Sugar-cane insects of the British West Indies) Dr. W. J. Hall, C.M.G., M.C., London (Hemiptera: Coccidao) Dr. D. Hille Ris La,mbers, Holland (Hemiptera: Aphididae) Professor Dr. A. cla Costa Lima, Hio de Janeiro (Sugar-cane insects of Brazil) Sir Guy A. K. Marshall, KC.M.G., F.R.S., London (Coleoptera: Curculionidae) Mr. R. W. MUllgomery, Brisbane (Sugar-cane insects of ) Dr. C. E. Pemberton, Honolulu (Sugar-cane insects of Hawaii) Dr. A. Reyno, Amsterdam (Sugar-ca.ne insects of Netherlands Guiana) lvir. L. C. Scaramuzza, Cuba (Sugar-cane insects of Cuba) Dr. B. P. Uvarov, C.M.G., F.R.S., London (Orthoptora) Mr. R. H. Van Zwaluwenbul'g, Honolulu (Coleoptera: Elateridao and Tenebrionidae as well as many genera in other orders; sugar-cano insccts of Hawaii) Mr. C. Vinson, Mauritius (Sugar-cane insects of Mauritius) Dr. J. Wille, Lima (Sugar-cane insects of Peru) Dr. G. N. Wolcott, Puerto Rico (Sugar-cane insects of Puerto Rico, Haiti, and the Dominican Republic). Mr. R. H. Van Zwaluweuburg, Entomologist of the Experiment Station of the Hawaiian Sugar Planters' Association, deserves very special mention for giving unsparingly of his time to answer innumerable enquiries, aside from his personal contributions to this List. It is also through his efforts, and those of Mr. R. Urata, of the Staff of tho same Experiment Station, that a translation was made available to tho writer of the indispensable eatalogue of the sugar-cane insects of, by Shuzo Takano and Masayuki Yanagihara, published (in Japanese) in 1939, which colltains nearly 250 separate records. It would be ungrl'tteful not to place on record the help that the writer has derived from a study of the magnificent work, recently published in two volumes, Die Plagen van den Cultunryewassen in indonesie., by Professor L. G. E. Kalshoven of Amsterdam, which because of its up-to-date nomenclature and the care given in citing references, has been of the utmost aid to an understanding of the sugar-cane insects of Java and adjacent territories. 3

12 At the Imperial College of Tropical Agriculture, Trinidad, Mr. R. G. Fennah has been particularly helpful in checking the large number of records (more than 200) of Fulgoroidea and Cercopidae, many of which were previously unidentified; he has also contributed several other records that would otherwise have escaped notice. Miss M. E. M. Price, Chief Librarian at the College, and her Stafl', have rendered much assistance in the search for literature. Neither must be forgotten the invaluable help given by Dr. 'V. J. Hall, C.M.G., Director, and Dr. T. H. C. Taylor, Assistant Director, of the Commonwealth Institute of Entomology, in dealing with the very considerable correspondence relating to this List, and by the Editorial and Library Staffs of the Institute, in particular Miss R. M. Davenport, for the care given to its production. To all of the above the writer acknowledges his indebtedness for their kind assistance and takes this opportunity to express his most sincere thanks. 4

13 PART.I SUGAR-CANE INS1WTH The first column gives the family, genus and species of tho insect feeding on sugar-cane, with SUOII a,lternative names, dnly cross-referenced, as have been llsed in the literature of sugar-cane eiltomoiogy; thcre m1itlos are not neoessarily i:lynonyms. The second column names the countries where the insect has beon recorded or o bserved on sugar-cane, but does not lleeesl-iarij:1l indicate the full geographioal distribution of the species. The third column records the para~ites fmel prechttol's by genera, 0111.'/. If a gen.us contaiur more than one species parasitic or predacious Oil it pilrticnlm' hol-lt, the lllunber ilo: given in bracket::;. The ]lf1ge reference::; to thu genora lilulor which tho specich are lii:lted in Part II may be obtained from the lmlex..

14 Nitidulidae Urophorus humeralir, E'. (probably) COLEOPTERA U.S.A. (Ga.) Bostrychidae Apate liwnacha, F. (jrancisca, :1".) Dinotic1'wl minut1l8, F. Enneadc81n1l8 forficula capcn8is, LeRne Cuba, Puerto Rico Indio. Mozambique (Lonren~ Marques) Anobiidae.X!lictinu.~ bra8ilicn8is, Pic,'ar. (Ll'(lentinus, Pic Argentina Elateridae AdeloccTa 8ctiycra. Bates (Lacon) Aeolorlcrma brach mrllta, Cando (A(lolu8 vittatus, l\fllts.) Acolus cinnrlylwiiwll8, BaiRd. (see 8imor1actylu,9) dorslll!", Say (see DI'!l8tCI'ius mdlill us, Say) elegan8, F. (see Dl'Cl8terius) viltatu8, Mats. (see Acolodemza brachmana, Cand.) Agonisch iu8 obscuripes, Gyl1. var. viuiger, Heyd. Agriotes jormosclnus, Mats. (see Platynychius) sacchm'i, Mats. taichuens-is, Mats. Agrypnus ju8cipcs, F. IfUistCI'wi, Pnsc. (see Part II) palitns, Cando Ga1'(liaphol'lls Mauritius. Has been recorded in sugar cane fields, and is probably a predat,ol' on Mololonthid larvae dr.mlstans, Mats. (see Paracardiophorus) j01'1nosan1l8, Mats. (see PlatYl!lIch-U.s) Gompsolacon j0l'1nosonu8, Bates (Lacon) Gonoderus (11iolloCl'ephliI18) a'l.lstralasiae, Boisd. bijoveatu8, Beau,. ex8111, Sharp liv'idu8, Deg. Forrnosc1.. Cuba, Puerto HieD Hawaii (immigrant) U.S.A. 6

15 Elateridae-continued Conodei'IlS (subgell, H etei'odel"l~8) albicails, Cand, amplicolli&, GyU, (laurentii, Guel',) as8us, Cando (see Lacon) cai'i'nsensis, Blackb, carilltltu,<, Blackb, humili,~, Erichs. (see Lacon) lateralis, Schwarz (see Lacon) ~FOl'lnOS!t U,S,A. (Fla,). Cuba lal/,rel/tii, Guer. (see amplicollis, Gyl!.) obutus. Cando G'repidomcnlw queensland'lells, Blr, Drasteriu.s' (Aeolus) elega'lls, F, West Indies rndlilllls, Say (dorsalis, Say) U.S.A, (Fla.) Helm'ot/eres (see Oonoderu8) Lacon (some of the species have been recorded as Hctcl'ode1'es) as:;i'!ls, Cund, (see also Part II) jol'mosanus, Bates (see Oompsolacon) humilis, Eriehs. Queonsland lateralis, Schwarz nmsculus, Cando (shirakii, Mats,) Formosu setiger, B(ttes (see Adelocera) shiraki'i, Mats, (see musculus, Cand.) strict~'colli8, Fairm, Fiji variabilis, Cando, Fiji variolu8, Cando Ludius suturalis, Mats. (Bee N eotrichoph01'u8) lv.legapenthes Cuba Ilt elanot'tl8 communis, GyU. U.S.A, (Fla.) tam8uyensis, Bates Ryul,yu Is" Jamaic(t Jl1onocrepidiu8 (see OonoderusJ N eotl'iohophorus sut1l1'ali8, Mats. (Ludiu..) OrthoBtethu8 infu8catu8, Germ, U.S,A. (Fla.) Paracar'diophol'1l8 devastan8, Mats, (Cardioplwru8) Platynychu8 jormosanu8, Mats. (AgrioteB, Oardioph01'U8 ) Sephilus jo)'mosan~ls, Schwarz Silesis mutabilis, Bates Simorlactylus cinnamomcus, Boisd. (Aeolus) Fiji, Hawaii (immigrant,) Buprestidae Aphanisticus consan(lltincu8, Ritsema krugeri, Ritsema Java Java Lycidae 1'honalmu8 bicolol', L. (rnilitaris, Dalrn,) 7

16 Tenebriollidae Daslls cf1l'pentariae, Blackb. (Gonoceplwlum) rleprc88us, F. (Gonocephalum, Opat7'um), scriatu8, Boisd. (Gonaccphal1tm) Hawaii (immigrant) Epyrill Caroline Is. Dystalica mackayens'is, Cart. Entochira lateral is, Bah. (see Eutochia) Eutochia (has been mis-spelled Euptochia) latemlis, Boh. (Entoehira; Holaniara picescens, Fairm.) Java, (Negro»), Guam (in rotten sugarcanes), Hawaii (immigrant,) Mauritius }Jnlla, Eric. Gonocephalum acutan.qulutn, Fairm. (see Opatrum) arlprm8um, Germ. llequatoriale, Blanch. bil'inealllm, \Vlk. em'penlariae, Blackb. (see Das1tu) depl'es8wn, I? (see D(18118) lwjjmannsey(ji, Stev. (MYRore) pabens, Marseul Formo~a se1'2'at1tln, Boisd. (see Dasus) simplex, F. tol'l'idurn, Champ. Holaniam picescens, Fairm. E1ttochia lateral'is, Bah.) Hopatrinus (see Opalrinlls) Opatrinu8 (Hopatrinus) uerncllatu8, Oliv. Opntrum (loutangulum, Fairm. (Gonocephnlnm) depl'es.wm. F. (see JJasus) : an early name for D. (Gona CB1Jhalwn) carpentariae Alleculidae (Cistelidae) D'imorphoeh1:Z1w pascoei, Mac]. Anthicidae.F'ormicomU8 caeruleipennis, Laferte impemtor, Laferte Meloidae Epicauta waterho'usei, Haag Rutenbul'g Melolonthidae* A ncistrosoma klu(ji, Curtis vitti(lej'u1n, El'ichg. Mauritius Queenslancl (see St. Lucia.Java. Guam Peru Argentina.. A number of species recorded in the literature as sugar-cane pests in nrc omii;ted on the advice of Mr. Mungomery, who points out that they have been found in collections of adult beetles freqnenting the same feeding trees as the pest species, and recorded in error. (See alba Rutelidae.) 8

17 Melolonthidae-continued Aneylonycha (Lachnosterna, Holo t1'icitia; see also lfl!'l:zotl'oglw) constrictor, Burro. J'uV(, jlackei, Brenske qcilenlwuscl'i. Brenske j'cllci'i:, Brenske horirilllna., Niijima & KinORhitlL leucophtlwlrna, IVied. mimjanaoana, Hronske sa. uter';, Moser SC1'l'uta, F. 'vz'cz,ua, Sharp A noplosia,gum rufurn. Chapin Apogonia bakeri, Moschl. destl'uctor, H. BOB proxima, Waterh. sauteri, Moser sp... 1.~Cl ica,java Forrnoso..laNa (Negros) Fonnosu lndi!t Java, Madagnscar Burma Cuba Java jormosae, Brenske (Autose1'ica) injlmw/a, Moser Autoscrica jol'mos(j(', Brenske (sce Ascl'ica.),inoan<lvilis, Brenske nigl'ol'uvra, Brenslw 2 spp. CCl'icesthis pnl'inosa., Dalm. Olenw1'U (Lachnostcma, Phyllophaga, Phytal'us) apicalis, Blanch. (inslllaris, Smyth) smithi, Arrow Onema1'achis (La chn08tcl" na, Phylloplw(Ja) ael'u(jino,m, Burm. Cuba anl1lis, BtlI'm. (vijavca/a, Jaeq.) Cuba (Lntiguae, Arrow Antigua In/oveata,,Jacq. (S00 (1n(lli8, Burm.) cl'cnaticollis, Blanch. Cuba (il:s8im.il'l:s, Ohevr. Cuba guan'i!!ana, Smyth Puerto Rico hagarrl1:, Blanch. ialll(licensis, Arrow neglecta, Blanch. patruelis, Chevr. pm toricen Si8, Smyt,h publ!l'ula, Jacq. 8utl~mli8, Chevr. vand'inei, Smyth Oochliotis mel(jlonthoiilcs, C:orst. Cuba, Puorto Rico Barbados, Mlluritius, (acci dental introduction from Barbados) Haiti,Jamaica Hait,i. Dominican Rop, St. Kitts Pucrto Rico Cuba Cuba Puerto Hieo 'l'unganyi ku o Campsomeri8 Oampsonwris Oa.mpsonwri8 (5), &olia (2) Gall!p80mCl'i8 Proscna lljyzin1l1ll PiJrojJlI01'1t8; UlIlJIjJsomeris (H), SGolin, 'l'ipkia Pl1l'aHitns I.)]'(dmhly UIO RllllW fls thll80 of 1)(11'10'l';''<','/I.8i" aud 'Vunrlinc'i Pt'ilorkm:rt P!JI'Op/t(JI'U8 py1'oph01"l18; Ptilor.ie,r,io, 'l'iplt:ia. ~, Pyrop/to'l'l18 ; O':t'lI1P8()IlW1'is (:l). lvlyzinwn (2); G'r!Jptolllci gen'in, Eutri~)oide8, 8"1' :0 plw(la P!J,'oplWI'U8; Oall!pSOmeri8 (a), IVlyzinwn (2); G'1'!JplIJ1/wi. genia, lcutrixoitic&

18 Melolonthidae--continlled Demzolepida albohil tum, Waterh. (LelJiriiota, Lepidodc1'Ina) Gymnogaster buphthalrna, Blanch.. (an early misidentification of Hhizotl'O{lU8 spp.) (Maul'it.iuR I HalJlonycha castanea, Mac!. (has been reo corded in error as HaJllonyx) dilatala, Lea Hetr;Tonyx Holotl'ichia (see Ancylonyr'ha) Hoplia maculifera, Motsch. Hoplochelus r/tizotl'ogoides, Blanch. Madagascar H yp08c/' iea 8l(omctallicu, Arrow Mauritius vin8oni, Arrow Mauritius Lall/mostemu (Ho[()tl'ich'ia) (seeancylonycha) Lacitrwsterna (l'h!llloplwga) (see UlelllOl'fl. Unemamchis, PhyUopl!aya) (Phylalus) (see Olcmom) Dachno8tcI'na (Rliizotl'ogus) (Bee Rhizotl'ogus) Lepi{liota albohil'la, Waterh. (see Dermolel'iria) blal1clianli, Dalla Torre caudcltn, Bluekb. Qneensland colisobrina, Gir. frenchi, Blnckh..froygatti, Macl. g/'ata, Blackb. (this was at ono time confused with tl'ic!lo8te1'/w) pruinosa, \Vied. rothei, Blackb. stigma, F. /,riclw8tel'lla, Lea (has been Qneollsland.Java confused wit.h grata, Bluokb.).LcpiriOr/O'lllfl (Ree, Jjl'i'il/oll'l)'idri) Leucoplwli8 il'l'ol'clla, Chevr. rorilla, F. Nriel'otrichia jol'lnosana, Moser Ncao,liwaI1:s, Blackb. flcwipcnnis, Macl. Phyllognatlms riionysius, F. silcnu8, F..Java Egypt Ph.ljllophagG (Dachno8terna) (see also Clelnora, Ommwmchis) antcnnata, L. bui'ineisteri, Lee. clypcata, Horn COn(/I"IW, Leo. U.S.A. U.S.A. U.S.A. (Flu.) U.S.A. 10 Agr!lpnu8; Oampsomeria (3), Scalia, 'l'risiloa.; Pl'omachus, Palpo8tollll1, Prosena, Huti Ua,."'tllTlIl ia, Sarcophaga Oampsomcl'is (2) Oa'lllpsomcris (a) Uampsomeri.' (:3) Caml'sonwris (:3); Plllpostoma, Rutilia Campsomeris (7), Scolia (2), Tiphia Camp8omel"is (2); Rutilia Palposloma Campsomcris (7), Scolia (:l), 'l'iphin ; Eulrixopilolis, Eutrixop8is

19 Melolonthidae-contimted PhyllopJUl.qn-continued c:rassib',r:~hn(jj Blanch. latijl'on.9, Lee. or spp. Phyt(1/U8 (~ce Ole1ilow) PSDudJwlophylln juljul'acea, BurID. Psilonychus Rhizotro{!us (Lachn,08terna)* gravis, Arrow (hus beon recorded in error as GymnoflC18te>' buphthalma, Blurwh.) pallens, Arrow RJlOpaea subnitfda, Arrow vestita, Arrow v'iticnsis, Fairm. Schizonycha vastatrix, Chiaramonte Semanoptel'U8 dep re88'1:'u8culus, Mac]. Serica u8.wnnen."j'i8, Bronske luy'ubri8, Brenske Stephanopholis philipp'i:nensis, Brenske U.S.A. U.S.A. (Fla.) Mexico Tangllnyilm Mauritius Mauritius Fiji Fiji Fiji Somalia Java Lacon; Oampsomel'is; Neoa Tatus ; Palpostoma, Rutilia Carnpsome"is Cmnpsome)'is Camp8omeri8, BeoUa Oonoderus; Oampsomeris, Scolia l'iphia Proscna Camp801l!C1'lR (G), Scolia (2) Rutelidae ** Adoretu$ caligino8lm, Burm. compl'e88u,~, Web. (appears to have been confused with A. versutu8 in Mauritius) Java Pro.scna jormosanll$, Ohau8 lui'idu8, Blanch. Camp.sOllWri8, 'l'i:phia mnuncuzt(s, Burll1. Mauritius, sinicu8, BUl'Il1.,, Hawaii (immigrant) tenuimacuzat1t8, Waterh. (mn- Japan,, Fiji brosus tenuimaculatu8, Waterh.) ~anbroslls, F. Java umbro8u8 tenuimuculatu8, Waterh. (see tenuirnaculatus) versulu8, Har. (see nato Madagascar. Maul'ij,ius, Oa.mpsomm in (~) under comprcssu8), Fiji AnomaZat ael'ea, Perty (seo Aprosterna untiqlla, elyl!.) ano(futtata, Burm. (JmnpsonlC'l'il1 (2) antiqua, Gyll. (see Aprosterna.) uustrazasiac, Blaekb. (seo Aprostcma antiqua, Gyl1.) badia, OhauR FOl'lllosa bengalens'is, Blanch. * The two Muuritian RIlecioA werd originully doscribecl under Rhizot.TOg'lt8. The compiler retains t,his generic pesition pending a revision of this and alliod gonera. ** See footnote under jj:felolontmdac (p. 8). t Dr. g. A. Chnpin informs the writor that Apro8tm'na and Euchlom, hore treated lls gonoi'ii (following Kalshoven, lugl), are usually regarded us subgenera of Anomula. II

20 Rutelidae-continuecl A nomala---cont'inued bihw'cns1b, Arrow caleeala, Chevr. caslaneoventris, Bates cupripe., Hope dimidiata, Hope (?) dussumieri, Blanch. erubescen8, Ohaus expansa, Butes humeralis, Burm. timbijera, OhullR nerissa, OhallB obsaleta, Blanch. orientalis, '\Vuterh. pallida, F. (see Aproslerna) pulchripes, Lansb. (see Euchlara viridis, var. pulchripes, Lansb.) quadricalcw'ata, Ohau8 sulcatula, Burm. lrachypyga, Bl1tes ustulata, Arrow t'cm:color, Gyll. (see Aprosterna) vi1'icus, P. (see Euchlora) xuntlloplenm, Arrow (see EuchlO1'a) Anoplognathus bo-l8rl'wuali, Boiad. J)alliclicollis, Blanoh. pan'ulub, W atel'h. (maslel'si, Mac!.) (GaUoodes) Aptosterna (AnDmala) antiqua, Gyl!. (aerea Perty, A. O1lstralasiae, Blackb.) pallida, P. v(l1'icolol', Gyll. Bolax fia?jolineata, Mannh. OaUoodes mastm'si, Mao!. (Bee Anoplognathu8 parvulu8, Waterh.) Euchlora (Anornala) viridis, 1". val'. pulch1'ipes, Lansb. xanthoplew'a, Arrow Mimela testaceoviddis, Blanch. Cetoniidae Cacochroa decorticata, Mac!. Oetonia rnanda1'ina, Weber (see Protaetia fusca, Hbst.) Prolactia fu8ca, Hbst. (Getonia manda dna, Weber) Dynastidae Aegopsi8 west7ljoodi, Thoms. Ali8sonotu1n C1'a8sum, Arl'cw imp"essicolle, Arrow pauper, BurID. piceurn, P..9irni/e, Arrow Cuba Java Hawaii (Oahu) (immigrant or accidental introduction from Japan), (?), Caroline Is. Mozambique (Lollren')o Marques) Queonsland Burma,,Java, Java, Java, Brazil (Rio) Javl1 Jl1va Ryukyu Is. Javl1 Java Venezuola Burma, FormoBI1, Burma,. Mauritiust (C. Vinson), Carnpsomeri., CampsDmeri,9 Campsomcris Carnpsomcri.9 (2), Tiphia Pr08cna (Jmnpsomeris, Tiphia (2) Gampsomeris, T'iphia Gampsomeri.. (3), Scolia (2); Rutilia ScDlia, Proscna Eutrixop8is Gampsomeri8 Garnpsomert's Gampsomeris

21 Dynastidae--co-ntinlled Chalepides barbatus, F. (DlJ8cinetu,~, E'lletheola, l'amchalepus) Puerto Rico, Leeward Is. Chalepu8 (RCC D!ISoinctlls) Cycloccplwl(l borcahs, An. (vill()sa, Bllrm.) U.S.A. eulmua, CllflJlin (signata, auot. Ilec F.) Cuba immaculata, Olk. (Ochrosidia) U.S.A. (Fla.) signata, F. British Guiana signata,,met. nee F. (sen cubana, Chapin) tridentata, F. Guadeloupe villosa, Burm. 11ec Blanch. (soe boreaus, An-.) DaS!lflnathu8 australis defeani, Mac!. D!fscinetu8 (Clzalepus, Parachalcpus) bm'batus, F. (see Chalepiclcs) birlentatlls, Burm, (Euethcola) duijiu8, Olivo (r/eminatus, F.) picipes, Burm. trachypygu8, Burm. 2 spp. (one of these has been recorded in error as bidcntatu8, Burm.) Elwtheola barbata, F. (see Chalepiclcs) birzentata, Burm. (see Dysc?:netus) humilis, Burm, (Ligyrus, Portal.gus) rugiceps, Lee. (Ligyrus) Goloja aegean, F. portcri, Hope p'llsilla, Arr. British Guiana British Guiuna Cuba, Puerto Rico U.S.A. Moxico Brazil U.S.A. (La.), Mexico PH Peru Peru Mexico (Morelos}t 8{mchez N., 1945) te1'sander, BUl'ITI. Mexico (Veracruz),!, H etcronljch1',~ licas, KIng Mozumbiqne, Natul Uoderes, Redt. Burma mol'atm', F, Java prwli1:, At,!'. (8acclwl'i, Paoli) Somalia pa1'1lmpunctatu"s, Burm. Egypt plebejus, Klug Madagascar saceila.?';, Paoli (Aeo paorii, Arr.) soncti lwlenae, Blanoh. New South Wales 8ublae1Jis, Fairm. t"lstis, Boh. Natal (apparently an error; BOO 111 etmnasies vulg1!jaou8) Homophileul'us (l'hilcu:/'us) quadritulwl'culat1ts, Beauv. (bajulus, Perty) 1sodon pnncticollis, Macl. L i01/1'us bituberculatu8, Beauv. (Tonuz1'll8) cunic'llljtb, F. (tumuloblls, Burm.) ebcil1l8, Deg. j08,'ator, Burm.. j08801', Latr. gibboslts, Deg. () British GniaIlEt QUOOllslund (F. Sarcophaga Scolia (2) ; Rutil'ia (2) Tiphia, Tiphiu 'i"i]jhia ; JUra:!.',2vleyup,winp8i,q, j.\i'icl'opittlialm/i. 8(l"colJhaga Brazil Cuba, Jamaica., Puerto Rico, GampSOrnC1'i8 St. Croix, St. Kitts, Nevis, Antigull Venezllola, British Guiana, Camp80I1Wr'i8, 'l' iph,:a Colombia Brazil Cuba U.S.A. (Ln.) 13

22 Dynastidae--continued Lig!ll'us-continued humil{s, Burm. (see Euetheola) maimon, Erichs. (Tmnarus) Peru I'llgiceps, Lee. (see Euetheola) sallad, Bates Mexico (Veracruz)t tumulo8us, Burrn. (see cuniculus, ]'.) zapeczmn, Burm. (error. see Strategu8 sal'peilon, Burm.) (Cuba) 111 etanastes vulg{t'agu8, Olliff (Pentodon australis, Blackb.). HUB been recorded in error us Hete1'onychu8 OchTosirZ,ia (see Cyclocephala) Oryctes monoceros, Olivo rhinoceros, L. tarandu8, OIiv. immaculata, Olivo Bp. Natal Pamchalepu8 barbatu8, F. (see ClwlepicZes) Pentodon australis, Blackb. (see ~Meta nasies vulyivclgus, Olliff) blmgalen8io, Arrow bispinijl'ons, Raitt. dispm', Burm. idiota, Hbst. Phileurus baiulu8, Perty (see Zanzibar, Burma, Indo China. Java, China, Korea, Mauritius. (Records from Madagascar appear to be erroneous) Egypt Persia H omnphileurus quadl'ituberculatu8, Beauv.) Podalyu8 hwnilis, Burm. (see Euetheola) sa1"pedon, Burro. (see Strategus) Podischmw aye nor, Olivo lwrni, Sterub. Stenocrates labomtor, F. StTategopsis sarpedu'tn, Burm. (see Strater/us sarperlon, Burm.) Stratcgu8 aia:r, Olivo (titanu8, allct.) aloeus. L. (has been confused with anachorctl18) anacliol'etu8, Bm'm. anaclwretu8, aud. nee Burm. (see aloeus, L.) barbiyel'lis, Chapin (titmzus, Guatemala, El Salvador, t Colombia, Venezuela Venezuela?Mexico (Veracruz)t, Brazil Cuba Trinidad, British Guiana, Colombia Cuba Ptilodexia, Rhamphinina Scolia Ca.mp8omeri8 (Trinidad) Jamaica(?), Dominican Rcp., auet.) Puerto Rico, St. Croix Camps01nel'is; Smcophaga oblollyu,s, Beauv. (qlladl ifo. veatll8. Burm.) Dominican Rep., Puerto Rico 8a1-pedon, Bllrm. (error, zapedwn) (Liyyru8, Porlal. fjus, Stratel/opsis) Cuba titanu8, auct. (see aiax, Oliv.) (Cuba) titanu8, auct. (sccljal'biyel"lls, (Jamaica, Puerto Rico, St. Chapin) Croix) validus, F. Argentina l'e11lnoithynchu8 baal, Reiche Egypt clypeatu8, Klug Natal H

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